Evolutionary Systematics. 5 2021, 193-261 | DOI 10.3897/evolsyst.5.63435
> PENSUFT.
EVOLUTIONARY SYSTEMATICS
An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH):
Part I. superfamily Papilionoidea
Reza Zahiri!:*, Vazrick Nazari?, Hossein Rajaei*, Martin Wiemers?, Maryam Fatahi', Matthias Seidel’, Thure Dalsgaard', Martin Husemann!
oF WN BP
http://zoobank. org/984E 15D8&-80E0-4B7D-A84F-92BB0AD4EA 73
Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany
Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., KLA 0C6, Ottawa, Ontario, Canada Independent Researcher, P.O. Box 45063, Ottawa, Ontario K2M 2Y1, Canada
Department Entomology, State Museum of Natural History Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany
Department of Systematics and Biogeography, Senckenberg Deutsches Entomologisches Institut, Eberswalder Str. 90, 15374 Miincheberg, Germany
Corresponding author: Reza Zahiri (reza.zahiri@inspection.gc.ca, reza.zahiri@gmail.com)
Academic editor: Danilo Harms # Received 22 January 2021 # Accepted 10 June 2021 # Published 20 August 2021
Abstract
We provide an updated catalogue of the type material of the lepidopteran superfamily Papilionoidea deposited in the Zoological Mu- seum of Hamburg (ZMH). We report 414 specimens labelled as “types” belonging to nine species (all of valid names), 74 subspecies (44 valid names and 30 synonyms), 59 invalid infrasubspecific names under the ICZN code, and 23 specimens of 16 “in litteris” (= unavailable) names. Out of the 414 specimens labelled as “types”, 171 specimens are primary types (8 holotypes/lectotypes and 163 syntypes) and 80 are secondary types; 120 specimens are infrasubspecific and hence invalid and are considered as “original specimens”; and 43 specimens are treated here as “Non-type” specimens (topotype). We present a full bibliography of the original
descriptions and illustrations for all of these taxa, aiming to provide a comprehensive taxonomic guide to this collection.
Key Words
Biodiversity, CeNak, Herbert Weidner, ZMH, digitization, Georg Warnecke, Hanan Bytinski-Salz
Introduction
Natural history collections are the main archives of biodiversity. They focus on collecting, maintaining and documenting natural specimens, a crucial task, especially in times of global biodiversity decline (Dirzo et al. 2014). The Zoological Museum of Hamburg (ZMH) holds large collections across all animal groups. With approximately five million specimens and several thousand primary types, the entomological collection is among the most important natural history archives in Germany. Yet, the collections are overall not well documented and updated type catalogues are largely lacking.
The last ZMH Lepidoptera catalogue was published by Herbert Weidner, a former curator of the collection,
in 1974 (Weidner 1974). Since the publication of this work more than 40 years ago, no further types have been documented. We initiated a comprehensive effort to doc- ument all of the type material preserved in the ZMH in English language (Sartori et al. 2016; Dey and Husemann 2018a, b; Harms and Dupérré 2018; Monod et al. 2019; Henningsen et al. 2020; Simoes et al. 2021; Zahiri et al. 2021). We performed a detailed inventory throughout the collection. As a part of this inventory, the entire collec- tion was thoroughly searched and types were transferred to a separate, specially protected type collection. All type specimens were compared with the original descriptions, photographed, databased and relabeled, if required. The original high-quality photographs are available online through the FUNDus! Portal of the University of Ham-
Copyright Reza Zahiri et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
194 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
burg (https://www.fundus.uni-hamburg.de/). The com- plete data of the type material is provided in two table formats: taxa format (Suppl. material 1: Table S1) and specimen format (Suppl. material 2: Table S2). Our first Lepidoptera catalogue (Zahiri et al. 2021) on the super- families Hepialoidea, Cossoidea and Zygaenoidea has resulted in an updated list of 259 as “type” labelled spec- imens belonging to 13 valid species, 34 subspecies (16 valid and 18 invalid names), and 36 names of invalid in- frasubspecific rank (based on the ICZN code).
Material and methods Photography
All images were taken with a custom-made digital im- aging system (DUN Inc.) with stacking capabilities. The images where either taken with a Canon EOS S5DSR with a 65 mm or 100 mm lens and stacked with Zerene Stacker (PMax algorithm) or with a Canon EOS 6D with a 55 mm or 65 mm lens and stacked with Helicon Focus 5.3.
Taxonomy
All taxonomic changes were made after studying the rele- vant original descriptions of the type material. Synonyms that have already been proposed are listed only if discussed and the type material was examined. Classification used herein follows van Nieukerken et al. (2011). The high- er taxonomy of the families Papilionidae, Hesperiidae, Pieridae, Lycaenidae, Riodinidae, and Nymphalidae, as well as the classification below the family level presented here, are based on a combination of recent comprehen- sive morphological and molecular studies (Paulus et al. 1996; Wahlberg et al. 2009; Heikkila et al. 2011; Kawa- hara et al. 2014), as well as the updated checklist of the European butterflies (Wiemers et al. 2018). Finally, like almost all major Lepidoptera databases and catalogues, we did not follow the principle of gender agreement in species names (Sommerer 2002). The recommendation not to follow the articles 31.2 and 34.2 of the ICZN code was adopted at the General Meeting of the Societas Euro- paea Lepidopterologica (SEL) in a resolution proposed at the 13" European Congress of Lepidopterology in Korsor (Denmark) on June 4, 2002 (van Nieukerken 2019).
Type material
Type localities are cited in their original spelling or with additional remarks (in square brackets), if required. All data was transcribed verbatim as appearing on the respec- tive labels. For unreadable data a question mark (?) was used. Double slash (//) separates data on different labels, a single slash (/) separates lines within each label, and semi-colon (;) separates different specimens. Additional comments are placed in square brackets.
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The ICZN (1999) terminologies (e.g., holotype, syn- type, lectotype, neotype, paratype, paralectotype, al- lotype, cotype etc.) are available at the ICZN website (http://iczn.org/content/Glossary) and ICZN articles cited in the text can be found below:
Infrasubspecific taxon — a taxon at lower rank than that of subspecies. The names of such taxa are not regu- lated by the Code.
Article 45.6.1. — A name is infrasubspecific if its author expressly gave it infrasubspecific rank, or if the con- tent of the work unambiguously reveals that the name was proposed for an infrasubspecific entity;
Article 45.6.2. — A name 1s deemed to be infrasubspe- cific if its author used one of the terms “aberration”, “ab.” or “morph”.
Article 45.6.3. — A name is deemed to be infrasubspecific if it was first published after 1960 and the author ex- pressly used one of the terms “variety” or “form” (in- cluding use of the terms “var.”, “forma”, “v.” and “f.”’):
Article 45.6.4. — A name is subspecific if first published before 1961 and its author expressly used one of the terms “variety” or “form” (including use of the terms “var.”, “forma”, “v.” and “f.”’), unless its author also
expressly gave it infrasubspecific rank, or the con-
tent of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, in which case it is infrasubspecific.
Type specimens deposited at the Zoological Museum Hamburg (ZMH)
Superfamily Papilionoidea Latreille, 1802
Papilionoidea (butterflies), currently containing seven families with approximately 18,800 described species worldwide (van Nieukerken et al. 2011), is one of the well-studied lepidopteran groups. The family Nymphali- dae (Brush-footed butterflies) with 6,152 described species is the most diverse taxonomic group followed by Lycaeni- dae (Blues and coppers) with 5,201 species, Hesperiidae (Skippers) with 4,113 species, Riodinidae (Metalmark butterflies) with 1,532 species, Pieridae (Whites or Yel- low-whites) with 1,164 species, Papilionidae (Swallotails) with 570 species and Hedylidae with only 36 species (van Nieukerken et al. 2011).
Family Papilionidae Latreille, 1802 Subfamily Parnassiinae Duponchel, 1835
1. tridentina Dannehl, 1925
Original combination. “Parnassius apollo L. (tridentina Dannehl)” Dannehl, 1925 Ent. Z. 39: 6.
Current combination. Parnassius apollo tridentina Dannehl, 1925.
Current status. Valid subspecies.
Evolutionary Systematics 5 2021, 193-261
Type material. Syntypes 24'¢ (ZMH 61579-61580) (Fig. 1). “Teriol. Mer. / Sum. alp. / Palon / 1.7.24 Ibor [?] / Dannehl” // “Cotype” // “1.7. 1924 / Mt. Palon / Stidtirol / Dannehl” // “f. tridentina Dhl.” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61579”; “10.6” // “Dannehl / Teriol. mer. / Alp. vai. / Mt. Palon / 1400 m.” // “Cotype” // “10.6 / Mt. Palon / Stidtirol / Dan- nehl” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61580”.
Type locality. Italy: “Lokalrasse aus den ostlichen Teilen der Trientiner (Sarcataler) Alpen; westl. Etschufer sudlich Trient. 1200—1600 m.”
2. mendolensis Dannehl, 1925
Original combination. “Parnassius apollo L. (mendo- lensis Dannehl).” Dannehl, 1925 Ent. Z. 39: 6.
Current combination. Parnassius apollo mendolen- sis Dannehl, 1925.
Current status. Valid subspecies.
Type material. Syntypes 1419 (ZMH 61581-61582) (Fig. 2). “mendolensis 2” // “Teriol mer. / Mendel / 24. juni / coll. F. Dannehl” // “Cotype” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61581”; “mendolensis 3” // “Teriol mer. / Mendel / 3.Juni.22 / coll. F. Dannehl” // “Cotype” // “Cotype” // “Mus. Altona comm / Eing Nr 3-65” // “ZMH 61582”.
Type locality. Italy: “Flegt auf den oberen der Men- delstrafe bei Bozen und in den meisten ostlichen Talein- schnitten des Mt. Roen, 500-1800 m.”
3. sevensis Kesenheimer, 1921
Original combination. “Parn. apollo Linné, nov. subsp. sevensis Ksh.” Kesenheimer, 1921 Ent. Z. 34: 63.
Current combination. Parnassius apollo sevensis Kesenheimer, 1921.
Current status. Junior subjective synonym of Par- nassius apollo meridionalis Pagenstecher, 1909.
Type material. Syntypes 1412 (ZMH 61583- 615584) (Fig. 3). “Cotype” // “apollo / sevensis / Cotype 9 / 19.7.14 / Aalfeld — Sternsee / H.W. Kesenheimer” // “Cotype” // “Sammlung / G. Warnecke / Eing Nr. 5, 1949” // “ZMH 61583”; “Sammlung / G. Warnecke / Eing Nr 51949” // “apollo sevensis / Cotype @ / 19.3.14 / Aalfeld - Sternsee / H.W. Kesenheimer” // “Cotype” // “Cotype” // “ZMH 61584”.
Type locality. France: “von Sewen im Masmiunstertal in den Vogesen”.
4. kricheldorffi Eisner, 1928
Original combination. “Parnassius apollo, forma kricheldorffi (m.)” Eisner, 1928 Int. Ent. Z. 22: 309.
Current combination. Parnassius apollo krichel- dorffi Eisner, 1928.
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Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 61585) (Fig. 4). “Asturien / Picos de Europa / 1800-2000 m. / 20. —28. VII.1928 / leg. A. Kricheldorff” // “v. Kricheldorfi Eisn.” // “Co-Type / e Collection / Bang Haas” // “Cotype” // “Parnassius V. / Kricheldorffi Es. / det. Eisner” // “Sam- mlung / G Warnecke / Eing Nr.5. 1949” // “ZMH 61585”.
Type locality. Spain: “Asturien, Picos de Europa, 1800—2000 m”.
Remarks. Eisner (1928) proposed this name as a form of P. apollo (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and the author expressly used the terms “variety” or “form”.
5. ab. praetexta Gelpke, 1929
Original combination. “Parnassius apollo L. f. valesi- acus ab. praetexta m.” Gelpke, 1929 Int. Ent. Z. 23: 123.
Current combination. Parnassius apollo valesiacus ab. praetexta Gelpke, 1929.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 12 (ZMH 61586) (Fig. 5). “Type / P. apollo L: valesiacus / Fruhst. / ab. praetexta Gelpke / I. E. Z. Guben 23 Jahrg 1929 / S. 122 [illegible]” // “13.7. 1927 / Lotschental / ob Wiler / Wallis 1550 m” // “Type” // “ab. praetexta Gelpke” // “Mus. Altona comm / Eing. Nr. 3-65 // ZMH 61586”.
Original locality. Switzerland: Swiss Alps. “oberhalb Wiler”.
Remarks. Gelpke (1929) proposed this name as an ab- erration of P. a. valesiacus Fruhstorfer, 1906. Therefore, as stated by articles 45.6.1 and 45.6.2 ICZN 1999) it is deemed to be an infrasubspecific name (if the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable.
6. ab. uniformis Stauder, 1922
Original combination. “Parnassius apollo ab. uniform- is” Stauder, 1922 Soc. Ent. 36: 10 (Note)
Current combination. Parnassius apollo ab. uni- formis Stauder, 1922.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 61587) (Fig. 6). “Unter-Italien / Stauder 1920” // “uni- formis Stdr / Type” // “ab. uniformis Stauder / Type” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61587”.
Original locality. Italy: “Unteritalien”.
Remarks. Stauder (1922) proposed this name as an aberration of P. apollo (Linnaeus, 1758). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.
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196 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
' Mus. Aitona comm Eing Ne 3-65
| Mus. Aitona comm’ Eing ‘Nre 3- 65°
z
ZMH - 61582
i ae i er | .
Mus. Aitena comm 2 Fing, Neo 3- 0d
Plate 1. 1. Parnassius apollo tridentina Dannehl, 1925; 2. Parnassius apollo mendolensis Dannehl, 1925; 3. Parnassius apollo sevensis Kesenheimer, 1921; 4. Parnassius apollo kricheldorffi Eisner, 1928; 5. Parnassius apollo valesiacus ab. praetexta Gelpke, 1929: 6. Parnassius apollo ab. uniformis Stauder, 1922. a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
7. f. bispupillata Turati, 1918
Original combination. “Parnassius apollo pumilus Stich. forma bispupillata” Turati, 1918 Atti. Soc. Ital. Sci. Nat. 57: 40.
Current combination. Parnassius apollo pumilus f. bispupillata Turati, 1918.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 14 (ZMH 61588) (Fig. 7). “Calabrien / [illegible], 1800 m / Stauder, 5.7.20” // “bispupillata” // “bispupillata” // “Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61588”.
Original locality. Italy: Calabria.
Remarks. Turati (1918) proposed this name as a form of P. a. pumilus Stichel, 1906. Therefore, as stated by arti- cle 45.6.1 (ICZN 1999) it is deemed to be an infrasubspe- cific name (if the author expressly gave it infrasubspecific rank) and is hence unavailable.
8. f. appendiculata Turati, 1918
Original combination. “Parnassius apollo pumilus Stich. forma appendiculata (f. nuova)” Turati, 1918 Atti. Soc. Ital. Sci. Nat. 57: 41.
Current combination. Parnassius apollo pumilus f. appendiculata Turati, 1918.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 1411 9 (ZMH 61589-61590) (Fig. 8). “Calabrien / Aspermonte, 1800 m / Stauder 5.7.1920” // “flavomaculata / appendiculata” // “flavomaculata / appendiculata” // “Sammlung / G. War- necke / Eing. Nr.5, 1949” // “Cotype” // “ZMH 61589”; “Calabrien / Aspromonte, 1800 m / Stauder 5.7.1920” // “nigricans / appendiculata” // “9 nigricans / appendicu- lata” // “Sammlung / G. Warnecke / Eing. Nr.5, 1949” // “Cotype” // “ZMH 61590”.
Original locality. Italy: Calabria.
Remarks. Turati (1918) proposed this name as a form of P. a. pumilus Stichel, 1906. Therefore, as stated by arti- cle 45.6.1 (ICZN 1999) it is deemed to be an infrasubspe- cific name (the author expressly gave it infrasubspecific rank) and is hence unavailable.
9. ab. lacrimans Marschner, 1911
Original combination. “Parnassius apollo L. ab. lacri- mans” Marschner, 1911 Dt. Ent. Z. Iris 25: 131.
Current combination. Parnassius apollo ab. lacri- mans Marschner, 1911.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 61591) (Fig. 9). “Schweizer Jura / Bozinger Berg / Juli
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1919/ecoll Marschner / coll. Dr. Gelpke / 1942” // “Type / Parn. apollo L. / f. lacrimans / Marschner / Collect. H. Marschner” // “ab. lacrimans Marschn.” // “Typen” // “Mus Altona comm. / Eing Nr. 3 05” // “ZMH 61591”. Original locality. Switzerland: “schweizer Jura”. Remarks. Marschner (1911) proposed this name as an aberration of P. apollo (Linnaeus, 1758). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.
10. oenipontanus Stauder & Felkel, 1921
Original combination. “Parnassius apollo oenipontanus, n. subsp.” Stauder & Felkel, 1921 Ent. Anz. 1: 123-124.
Current combination. Parnassius apollo oeniponta- nus Stauder & Felkel, 1921.
Current status. Valid subspecies.
Type material. Syntypes 104.4 (ZMH 61592-61601) (Fig. 10). “el. / 9./6.1924 / [illegible] / Innsbruck” // “Coll. Janiter / Leihgabe der / Jungiusgesellschaft / Eing Nr. 13, 1958” // “ZMH 61592”; “Type” // “Innsbruck / 10.6.1913 / J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61593”; “Type” // “Innsbruck Stilltal” // “Silltal /7.6.1921 / J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlu- ng / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61594”; “Cotype” // “oenipontanus / Stauder — Felkel / Innsbruk” // “Silltal / 23.5.1921 / J. Felkel” // “Coll. G. Warnecke / Eing. Nr. 11,1983” // “ZMH 61595”; “Type” // “Silltal / 19.5.1921 / J. Felkel” / “oenipontanus / Stauder — Felkel” /“Tnnsbruck / Silltal” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61596”; “Cotype” // “Innsbruck / gebirge / 21/6/1914 / J. Felkel leg.” // “oenipontanus Stder / Cotype” // “Sammlu- ng / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61597”; “Type” // Innsbruck / Silltal” // “Silltal / 19.5.1921 / J. Felkel” // “oenipontanus Stauder / Cotype” //“Summtng / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61598”; “Type” // “H. Stauder / Nordtirol / Patsch / 11/6/1920” // “oeni- pontanus Stauder / Cotype” // “Sammlung / G. Warnecke /Eing. Nr. 5,1949” // “ZMH 61599”; “Type” // Innsbruck / 2/6/1914 /J. Felkel leg.” // “subsp. oenipontanus / Stauder / Cotype” // “Summing / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61600”; “Type” // “Innsbruck / 23.6.1913 / J. Felkel” // “oenipontanus Stder / Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5,1949” // “ZMH 61601”.
Type locality. Austria: “Silltal von Wilten bei Inns- bruck bis Patsch, Tirol” [Sill Valley of Wilten near Inns- bruck to Patsch, Tirol].
11. serpentinicus Mayer, 1925
Original combination. “Parn. apollo vy. serpentinicus, (Rbl. i. 1.) Mayer, subspecies nova” Z. des Oster. Ent.- Ver. 10: 2-4.
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Current combination. Parnassius apollo serpentini- cus Mayer, 1925.
Current status. Valid subspecies.
Type material. Syntypes 24.4 (ZMH 824705-824706) (Fig. 11). “Cotype” // “Burgland / Rotlschlg. / Juli 24 / Mayer” // “Sammlung / G. Warnecke / Eing. Nr 5 , 1949” /1 “ZMH 824705”; “Cotype” // “Burgland / Stuben / Juli 24 /L. Mayer” // “Serpentinicus Mayer” // “Sammlung / G. Warnecke / Eing. Nr 5 , 1949” // “ZMH 824706”.
Type locality. Austria: “Auf einem Serpentingebirge an der ungarischen Grenze” [on a Serpentin mountain at the Hungarian Border].
12. ab. marschneri Bryk, 1911
Original combination. “Parnassius apollo ab. mar- schneri” Bryk, 1911 Int. Ent. Z. 5: 161.
Current combination. Parnassius apollo ab. mar- schneri Bryk, 1911.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 824719) (Fig. 12). “Nordeuropa / Stockholm / Umb. / coll. Dr. Gelpke” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “Type” //“Abgebildet / Bryk” // “Internat. / Entomolog. / Zeitschrift /Guben / V. Jahrg. No. 20” // “ZMH 824719”.
Original locality. Not indicated. Probably Stockholm [Sweden].
Remarks. Bryk (1911) proposed this name as an ab- erration of P. apollo Linnaeus, 1758, therefore, as stated by article 45.6.2 ICZN 1999); it is deemed to be an in- frasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
13. wenzeli Bryk, 1921
Original combination. “Parnassius apollo v. Wenzeli” Bryk, 1921 Ent. Tidskr. 42: 117.
Current combination. Parnassius apollo wenzeli Bryk, 1921.
Current status. Valid subspecies.
Type material. Synype 14 (ZMH 824721) (Fig. 13). “Cotype” // “Silltal / 19.5.1921 / J. Felkel” // “Coll. G. Warnecke / Eing. Nr. 11 1963” // “ZMH 824721”.
Type locality. “Innsbruck (Patsch)”.
Remarks. Bryk (1921) proposed this name as a va- riety of P. apollo Linnaeus, 1758. According to article 45.6.4 (ICZN 1999), it is subspecific if first published be- fore 1961 and author expressly used the terms “variety” or “form”.
14. gabrieli Bryk, 1934
Original combination. “P. nomion ssp. gabrieli (subsp. nova)” Bryk, 1934 Parnassiana 3: 28.
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Current combination. Parnassius nomion gabrieli Bryk, 1934.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 61602) (Fig. 14). “Co-Type / Collection / Kotzsch” // “Kukunor / Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Par- nassius / nomion / gabrieli 3 / Bryk” // “Coll. G. War- necke / Eing. Nr. 11 1963” // ZMH 61602”.
Type _ locality. China: “Burchan-Buddha-Kette” [ Burchan-Buddha-Chain].
15. chingamensis Bryk & Eisner, 1932
Original combination. “P. nomion F.d.W. chingamensis subsp. nova” Bryk & Eisner, 1932 Parnassiana 2: 26.
Current combination. Parnassius nomion chin- gamensis Bryk & Eisner, 1932.
Current status. Valid subspecies.
Type material. Syntypes 2¢'3' (ZMH 61604-61605) (Fig. 15). “Parnassius / chinganen- / sis 3 / C. B.-L / Chin- gan — Feb. / Ost- Mongolei / lec 7.” // “Coll. G. Warnecke / Eing. Nr. 11. 1963” //“ZMH 61604”; “Mandschur.sept.occ. / Buchalu / Chingan mont. sept. / 1100 m. Juli” // “chingan- ensis’ // “Staudinger / Bang-Haas / Blasewitz” // “Public. Bryk & Eisner / Parnass. I, 2, 1932, 26 / als 1” // “Sam- mlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 616035”.
Type locality. China: “Buchatu, Gudschal-Tal, Chin- gan mont. sept. 100 m.” [Buchatu, Gudschal-Valley, Chingan mont. sept. 100 m].
Remarks. This appears to be an erroneous original spelling, which was corrected by Eisner (1935) to “chin- ganensis”. However, Ackery (1973) treats this emenda- tion as an unjustified emendation according to ICZN arti- cle 33.2.3 (ICZN 1999) and not as a justified emendation.
16. tsinlingensis Bryk & Eisner, 1932
Original combination. “P. nomion F.d.W. tsinglingensis subsp. nova” Bryk & Eisner, 1932 Parnassiana 2: 26,
Current combination. Parnassius nomion tsinlin- gensis Bryk & Eisner, 1932.
Current status. Valid subspecies.
Type material. Syntype 1¢ (ZMH 61624) (Fig. 16). “Prov. Schensi / Tapaischan [?] / Tsinling schan / 3000 m 20.6.” // “Co-Type / e Collection / Bang Haas” // “subsp. tsinglingensis” // “Public. Bryk t. Eisner / Parnass. II, 2, 1932 / p. 26” // Sammlung / G. Warnecke / Eing. Nr.5, 1949” // “ZMH 61624”.
Type locality. China: “Schensi, Tapaischan, Tsinling- shan, 3000 m. VI.; Tsingschui, Liu-pinschan, 2000 m.”
17. nominulus Staudinger, 1894
Original combination. “Parnassius Nomion var. No- minulus” Staudinger, 1894 Dt. Ent. Z. Iris 7(2): 241.
Evolutionary Systematics 5 2021, 193-261 199
[zm » 61588
thus Altona com: fcing NRPS 0B :
ZMH - 61591
Nordeuropa 7 in tlng- coll. Dr. Gel pke
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Plate 2. 7. Parnassius apollo pumilus f. bispupillata Turati, 1918; 8. Parnssius apollo pumilus f. appendiculata Turati, 1918; 9. Par- nassius apollo ab. lacrimans Marschner, 1911; 10. Parnassius apollo oenipontanus Stauder & Felkel, 1921; 11. Parnassius apollo serpentinicus Mayer, 1925 12. Parnassius apollo ab. marschneri Bryk, 1911. a. Dorsal view, b. ventral view.
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200
Current combination. Parnassius nomion nominu- lus Staudinger, 1894.
Current status. Valid subspecies.
Type material. Type? 1419 (ZMH 824717-824718) (Fig. 17). “Irkutsk / Sibirien / A.M. Schmidt / Inh. M. Gottmann” // “Parn. nomion Hb. / Typus?” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 824717”; “Ken- tei-Geb.” // “Gebr. Dorries leg. 1893 / Erworb. e. coll. / Dorries sen. 1917” // “Nomion typ., [illegible]” // “Sig-G Warnecke / Eing. Nr 5 1949” // “Katalog Nr. / L. 604” // “ZMH 824718”.
Type locality. Russia: “Ost- Sajan -Gebiet (stidwestlich von Irkutsk, an der Grenze der chinesischen Mongolei gelegen)” [East-Sajan region (southwest of Irkutsk, at the border of Chinese Mongolia)].
Remarks. Staudinger (1894) proposed this name as a variety of P. nomion Fischer & Waldheim, 1823. According to article 45.6.4 ICZN 1999), it is subspecific if first pub- lished before 1961 and author expressly used the terms “va- riety” or “form”. We believe that these two specimens be- long to ssp. nominulus Staudinger, 1894. The year on one of the labels (1893) supports this. The question mark on the la- bel (“Typus?’’) shows uncertainty about whether it is a type. In the original description, Staudinger says: “... bemerke ich, dass die kleineren Stticke aus dem Kentei-Gebirge den grosseren var. Nominulus fast gleichkommen.“ [...I realize that the smaller specimens from the Kentei-Mountains are very similar to the larger var. nominulus].
18. frivaldszkyi Bang-Haas, 1928
Original combination. “Parnassius szechenyii frivaldszkyi O. B.- Haas, subsp. nov.” Bang-Haas, 1928 Ent. Z. 42: 60.
Current combination. Parnassius szechenyii frivaldszkyi Bang-Haas, 1928.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 61603) (Fig. 18). “Kansu occ. / Liang-tschou/ Richthofen mont. mar. /2500 m Juli” // “Co-Type / Haas” // “129” // “szechenyii / frivaldsz- ky?’ // “Mus Altona / Eing Nr 9-64” // “ZMH 61603”.
Type locality. “China sept. occ.: Kansu, Richthofen mont., stidwestl. Uang-tschou”.
19. ab. privatus Kotzsch, 1936
Original combination. “Parnassius autocrator Avinoft ab. privatus” Kotzsch, 1936 Ent. Rdsch. 54: 44.
Current combination. Parnassius autocrator ab. privatus Kotzsch, 1936.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 14 (ZMH 61606) (Fig. 19). “Hindukusch / Chodja-Mahomed / Gerollzone / 3800— 4000 m 25.VII.-10. VIII. / leg. H. & E. Kotzsch” //“Co-Type /e Collection / Kotzsch” // “Mus Altona / Eing 9-64” // “ZMH 61606”.
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Original locality. Afghanistan: “Hindukusch, Chod- ja-Mahomed, Ger6llzone, 3800— 4000 m.”
Remarks. Kotzsch (1936) proposed this name as an aberration of P. autocrator Avinoff, 1913. Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph’) and is hence unavailable.
20. lictor Bryk & Eisner, 1937
Original combination. “P. or/eans Oberth. subsp. /ictor (subsp. nova)” Bryk & Eisner, 1937 Parnassiana 4: 58.
Current combination. Parnassius orleans lictor Bryk & Eisner, 1937.
Current status. Valid subspecies.
Type material. Synypes 14112 (ZMH 61607-61608) (Fig. 20). “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / orleans / lictor 3 / Br. u. Ets.” // “Co-Type / e Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61607”; “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / orleans / lictor 2 / Br. u. Eis.” // “Co-Type / e Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61608”.
Type locality. China: Kukunor, “Burchan-Buddha, Nomohun Pa”.
21. buddha Bryk & Eisner, 1937
Original combination. “P. mercurius subsp. buddha (subsp. nova)” Bryk & Eisner, 1937 Parnassiana 4: 58.
Current combination. Parnassius jacquemontii buddha Bryk & Eisner, 1937.
Current status. Valid subspecies.
Type material. Syntypes 1412 (ZMH 61619- 61620) (Fig. 21). “Co-Type / e Collection / Kotzsch” // “Kukunor/ Geb. Burchan Buddha / Nomohun Pass / 4000 m Juli” // “Parnassius / mercurius | buddha & / Br. U. Eis.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61619”; “Kukunor/ Geb. Burchan Buddha / No- mohun Pass / 4000 m Juli” // “Co-Type / e Collection / Kotzsch” // “Parnassius / mercurius | buddha & / Br. U. Eis.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61620”.
Type locality. China: “Burchan-Buddha-Kette, No- mohun PaB, 4000 m.”
Remarks. Parnassius mercurius Grum-Grshimailo, 1891 is currently considered a subspecies of P. jacque- montii Boisduval, 1836.
22. ab. tripicta Sheljuzhko, 1928
Original combination. “Parnassius bremeri amgunen- sis’ 7. ab. tripicta (nov.)” Sheljuzhko, 1928 Mitt. Minch. Ent.Ges eis. 7;
Evolutionary Systematics 5 2021, 193-261 201
Kukuner Geb.Burchan Buddha Pass
CoH. 6, Warnecke Eing. Nr. 17 1963
zm : 61624
oy “Kansu eco. :
nahi ic! fen mont, mar. 2500 mull
Gebr. Dérries leg.1873 Erworb. e. coll. DGrries sen. 1917 _
“Mus Altona sepa gente
‘Katalog Nr. | CON
ZMH 824718)
Plate 3. 13. Parnassius apollo wenzeli Bryk, 1921; 14. Parnassius nomion gabrieli Bryk, 1934; 15. Parnassius nomion chingamen- sis Bryk & Eisner, 1932; 16. Parnassius nomion tsinlingensis Bryk & Eisner, 1932; 17. Parnassius nomion nominulus Staudinger, 1894; 18. Parnassius szechenyii frivaldszkyi Bang-Haas, 1928. a. Dorsal view, b. ventral view.
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202 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Current combination. Parnassius bremeri amgun- ensis ab. tripicta Sheljuzhko, 1928.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 14 (ZMH 61621) (Fig. 22). “Mandschur sept. occ. / Buchalu / Chin- gan mont. Sept. / 1100 m. Juli” // “Cotype” // “Co-Type / e collection / Bang-Haas” // “Bremeri solonensis / ab. tripicta u. flavicans” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 61621”.
Original locality. Not indicated. Probably “China, Chingan mont sept., Buchalu.”
Remarks. Sheljuzhko (1928) proposed this name as an aberration of P. b. amgunensis Sheljuzhko, 1928. Therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and 1s hence unavailable.
23. melaniticus Bang-Haas, 1915
Original combination. “Parnassius actius melaniticus O. B.-H., nov. var.,” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 7A 2A.
Current combination. Parnassius actius melaniticus Bang-Haas, 1915.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 833442) (Fig. 23). “Co. Type” // “3 P. actius /melaniticus” //“Kutscha mont. / Thianschan or / Juli 3500 m.” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 833442”.
Type locality. China: “Tianschan or. mer., Kour- gak Taon, Kuldscha Gebirge, nordwestlicher Kara- schahr”.
Remarks. Bang-Haas (1915) proposed this name as a variety of P. actius (Eversmann, 1843). According to article 45.6.4 (ICZN 1999), it is subspecific if first pub- lished before 1961 and author expressly used the terms “variety” or “form”.
24. wernickei Kotzsch, 1936
Original combination. “Parnassius charltonius Gray subsp. wernickei subsp. nova.” Kotzsch, 1936 Parnassi- ana 4: 6.
Current combination. Parnassius charltonius wer- nickei Kotzsch, 1936.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 61622) (Fig. 24). “Nord-Ost-Hindukusch / Nuksan-Pass Nordseite / Al- penwiesenzone / 3500-3800 m Mitte Juli / leg. H. & E. Kotzsch” // “Co-Type / e Collection / Kotzsch” // “983” // “Sig. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 61622”,
Type locality. Afghanistan: “Ost-Hindukusch: Nuk- san-PaB, Alpenwiesenzone 3500 bis 4000 m.”.
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25. inopinatus Kotzsch, 1940
Original combination. “Parnassius inopinatus” Kot- zsch, 1940 Ent. Z. 54: 17.
Current combination. Parnassius inopinatus Kot- zsch, 1940.
Current status. Valid species.
Type material. Syntypes 6¢'¢' (ZMH 61609-61612, ZMH 824997-824998) (Fig. 25). “Nordwest-Afgani- stan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 20.IV.-15.VH. / Exp. Wernicke” // “Parnassius / inopi- natus | f Kotzsch” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61609”; “Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesen- zone / 2800-3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61610”; “Nordwest-A fgani- stan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 20.1V.-15. VII. / Exp. Wernicke” // “Co-Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11 1963” // “ZMH 61611“; “Nordwest-Afganistan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Parnassius / inopinatus /m Kotzsch” //“Co- Type / Collection / Kotzsch” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61612”; “Nordwest-Afghanistan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 20.IV.- 15. VII. / Exp. Wernicke” // “Co-Type / e Collection / Kot- zsch” // “981” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824997”; “Nordwest-A fghanistan / Firuskuhi-Mont / Alpenwiesenzone / 2800-3000 m 20.IV.-15.VII. / Exp. Wernicke” // “Co-Type / e Collection / Kotzsch” // “980” //“Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824998”.
Type locality. “Nordwest-Afghanistan, Firuskuhi-Kette, 2800-3000 m.”.
26. gigas Kotzsch, 1932
Original combination. “Parnassius imperator gigas ssp. n.” Kotzsch, 1932 Ent. Z. 45: 267.
Current combination. Parnassius imperator gigas Kotzsch, 1932.
Current status. Valid subspecies.
Type material. Syntypes 2¢'3' (ZMH 61613-61614) (Fig. 26). “Sincheng / Ost-Sikingshan Gebg. / Juli 2800 m.” // “Parnassius / imperator / gigas f / Her.” // “Cotype” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61613”; “Sincheng / Ost-Sikingshan Gebg. / Juli 2800 m.” // “Par- nassius / imperator / gigas m / Her.” // “Cotype” // “Coll. G. Warnecke / Eing. Nr. 11, 1963” // “ZMH 61614”.
Type locality. China: “vom Ostabhange des Siking- shan Gebirges — Sincheng etwa 2800 m.” [Eastern slopes of the Sikingshan Mountains — Sincheng, ca. 2800 m].
27. clorinda Kolar & Querci, 1937
Original combination. “Parnassius mnemosyne L. V. clorinda” Kolar & Querci, 1937 Lambillionea 5: 99.
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Plate 4. 19. Parnassius autocrator ab. privatus Kotzsch, 1936; 20. Parnassius orleans lictor Bryk & Eisner, 1937; 21. Parnassius Jacquemontii buddha Bryk & Eisner, 1937; 22. Parnassius bremeri amgunensis ab. tripicta Sheljuzhko, 1928; 23. Parnassius actius melaniticus Bang-Haas, 1915; 24. Parnassius charltonius wernickei Kotzsch, 1936. a. Dorsal view, b. ventral view.
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204 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Current combination. Parnassius mnemosyne clo- rinda Kolar & Querci, 1937.
Current status. Valid subspecies.
Type material. Syntypes 2¢'¢ (ZMH 824701-824702) (Fig. 27). “Type” // “Macedonia / Olympus, 4500’ / June 16, 1935 / Rome” // “Balkan / coll. Dr. Gelpke” // “Mus Altona / Eing. Nr. 9 — 64” // “ZMH 824701”; “Typen” // “Macedonia / Olympus, 4500’ / June 19, 1935 / Romei” // “Balkan / coll. Dr. Gelpke” // “v. clorinda Querci” // “P. Mnemosyne / - Clorinda Querci / Type abgebildet / in Lambillionea” // “Type” // “Mus Altona / Eing Nr 9-64” /1 “ZMH 824702”.
Type locality. Macedonia: Olympus.
Remarks. Kolar and Querci (1937) proposed this name as a variety of P. mnemosyne L. According to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “variety” or “form”.
28. rjabovi Sheljuzhko, 1935
Original combination. “Parnassius mnemosyne rjabo- vi” Sheljuzhko, 1935 Zs. Ost. Ent. Ver. 20: 22.
Current combination. Parnassius mnemosyne rja- bovi Sheljuzhko, 1935.
Current status. Junior subjective synonym of Par- nassius mnemosyne caucasia Verity, 1911.
Original material. Labelled as “Autotypes” 1319 (ZMH 824703-824704) (Fig. 28). “autotypus 9 / rja- bovi Shel.” // “Kleinasien / Armenia / Daratshitshag / 7.VII.1939 / B. Tkatshukov leg. / coll. Dr. Gelpke” // “Mus Altona / Eing Nr 9-64” // “ZMH 824703”; “auto- typus 3 / rjabovi Shel.” // “Kleinasien / Armenia / Da- ratshitshag / 7.VII.1939 / B. Tkatshukov leg. / coll. Dr. Gelpke” // “v. rjabovi Shelj.” // “Mus Altona / Eing Nr 9-64” // “ZMH 824704”.
Original locality. Armenia: Daratshitshag.
Remarks. The year on the specimen labels (1939) in- dicate specimens were collected four years after descrip- tion (1935) from the same location (i.e., Armenia: Darat- shitshag) the original holotype material was collected. As a result, the label of the specimens treated as “autotypus” by Warnecke 1s erroneous. Autotypes do not exist in tax- onomy. The specimen collected from the same geograph- ical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these two specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
29. comitis Bryk, 1912
Original combination. “Parnassius mnemosyne vat. co- mitis” Bryk, 1912 Soc. Ent. 27: 39.
Current combination. Parnassius mnemosyne comi- tis Bryk, 1912.
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Current status. Junior subjective synonym of nomi- notypical Parnassius mnemosyne (Linnaeus, 1758).
Original material. Labelled as “Cotypen” 14 (ZMH 824707) (Fig. 29). “817” // “Italia central. / Majella / 21.VII.1925 / coll. F Dannehl” // “Italien / coll. Dr. Gelp- ke” // “Cotypen” // “v. majellensis Dhl.” // “v. comitis Bryk.” // “Mus. Altona comm. / Eing. Nr. 3 65” // “ZMH 824707”.
Original locality. Italy: “aus der Majella Gruppe”.
Remarks. Bryk (1912) proposed this name as a vari- ety of P. mnemosyne (Linnaeus, 1758). According to arti- cle 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the terms “vari- ety” or “form”. The year on the specimen label (1925) in- dicates specimen was collected 13 years after description (1912) from the same location (i.e., Majella Mountains, Italy) the original holotype material was collected. As a result, the label of the specimen treated as “Cotypen” is erroneous. The specimen collected from the same geo- graphical location from which the type specimen was col- lected is called topotype. According to ICZN (1999), a to- potype has no formal standing and 1s not regulated by the Code. Therefore, this specimen is erroneously labelled as type, should be annotated as “Not a Type” and is treated here as “Non-type” specimen.
30. gigantea Staudinger, 1886
Original combination. “Parn/[assius] mnemosyne L. vat. gigantea” Staudinger, 1886 Stett. Ent. Ztg. 47 (4-6): 197.
Current combination. Parnassius mnemosyne gi- gantea Staudinger, 1886.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 824713) (Fig. 30). “vy. gigantea / 1/86 3 / Tary. / Ob. Turk. / “Warnecke” // “Tary Ob. Turkestan” // “<4 Parn. Mnemosyne | v. gi- gantea / E.W. BETH / Hamburg. den Haag” // “Coll. H. Rodinger / Eing. Nr. 30, 1952 / Katalog Nr. L / 6133” // “ZMH 824713”.
Type locality. Russian Turkestan.
Remarks. Staudinger (1886) proposed this name as a variety of P. mnemosyne (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first pub- lished before 1961 and author expressly used the terms “variety” or “form”.
31. beicki Bryk & Eisner, 1932
Original combination. “P/arnassius] epaphus beicki subsp. nova (O. Bang-Haas 1. |.)” Bryk & Eisner, 1932 Parnassiana 2: 25, 42.
Current combination. Parnassius epaphus beicki Bryk & Eisner, 1932.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 824714) (Fig. 31). “Kansu sept occ / Hsining / Nanchan mont. / Tatung /
Evolutionary Systematics 5 2021, 193-261 205
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Plate 5. 25. Parnassius inopinatus Kotzsch, 1940; 26. Parnassius imperator gigas Kotzsch, 1932; 27. Parnassius mnemosyne clo- rinda Kolar & Querci, 1937; 28. Parnassius mnemosyne rjabovi Sheljuzhko, 1935; 29. Parnassius mnemosyne comitis Bryk, 1912; 30. Parnassius mnemosyne gigantea Staudinger, 1886. a. Dorsal view, b. ventral view.
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206 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Nadelholzzone / Jiili 2000 m.” // “Co-Type / e Collection Bang-Haas” // “Ep. subsp. Beiki” // “public. v. Bryk Ut. Eisner / Parnass. II, 2, 1932 p. 25” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 824714”. Type locality. China: “Std Tatung‘sche Berge, Kansu, 3000 m* [Southern Tatung Mountains, Kansu, 3000 m]. Remarks. Also misspelled beiki in literature.
32. ab. rubropicta Bang-Haas, 1915
Original combination. “Parnassius epaphus ab. ru- bropicta” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 95.
Current combination. Parnassius epaphus ab. ru- bropicta Bang-Haas, 1915.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Co-Type” 14 (ZMH 824715) (Fig. 32). “Karakorum mont. / Schahidulla / Chines Tatarei / 4000 m. Ende Juni” // “Co-Type / e Col- lection / Bang-Haas” // “subtilis / rubropicata” // “S\g. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824715”.
Original locality. Not indicated. Probably China.
Remarks. Bang-Haas (1915) proposed this name as an aberration of P. epaphus Oberthtr, 1879. Therefore, as stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
33. candidatus Bang-Haas, 1915
Original combination. “Parnassius delphius var. candi- datus” Bang-Haas, 1915 Dt. Ent. Z. Iris 29 (2/3): 158.
Current combination. Parnassius delphius candida- tus Bang-Haas, 1915.
Current status. Junior subjective synonym of the nominotypical Parnassius delphius Eversmann, 1843.
Type material. Syntype 14 (ZMH 824716) (Fig. 33). “Chin. Turkestan / Thianschan mont. or. / Juldus Tal / 2500 m. Juli” // “Co-Type / O. Bang-Haas” // “delphius — caudi- | datus Cotype” // “Slg. Mus. Altona / Eing. nr. 6/1978” // “ZMH 824716”.
Type locality. “Tianschan or.; Juldus (Kuldsca, Ill, Issykkul)”.
Remarks. Bang-Haas (1915) proposed this name as a variety of P. delphius Eversmann, 1843. According to article 45.6.4 (ICZN 1999), it is subspecific if first pub- lished before 1961 and author expressly used the terms “variety” or “form”.
34. eva Kotzsch, 1936
Original combination. “Parnassius delphius Eversm. subsp. eva subsp. nova” Kotzsch, 1936 Parnassiana 4: 5. Current combination. Parnassius jacobsoni eva Kotzsch, 1936. Current status. Valid subspecies.
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Type material. Syntypes 2¢¢ (ZMH 824723- 824724) (Fig. 34). “Hindukusch / Chodja-Mahomed / Gerollzone / 3800.4000 m 25.VII.-10.VIII. / leg. H. & E. Kotzsch” // “Co-Type / e Collection Kotzsch” // “O77” // “Sig. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824723”: “Hindukusch / Chodja-Mahomed / Gerollzone / 3800.4000 m 25. VII.-10. VHT. / leg. H. & E. Kotzsch” // “Co-Type / e Collection Kotzsch” // “978” // “Slg. Mus. Altona / Eing. Nr. 6/1978” // “ZMH 824724”.
Type locality. “Hindukusch, Chodja-Mahomed” [Af- ghanistan, Hindukusch, Khoja Mohammad Mts].
Unpublished, unavailable names
35. ab. minor Frank
Original combination. Parnassius mnemosyne hart- manni ab. minor Frank.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 1¢ (ZMH 824708) (Fig. 35). “Nordl. Kalkalpen / Inntaler Berge / Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // “Mus. Altona comm. / Eing. Nr. 3-65” // “Type” // “ab. minor Frank” // “Mus. Altona comm. / Eing Nr. 3 65” // “ZMH 824708”.
Original locality. Nordl. Kalkalpen [Austria].
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hart- manni Standfuss, 1888.
36. ab. nigra Frank
Original combination. Parnassius mnemosyne hart- manni ab. nigra Frank.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 1¢ (ZMH 824709) (Fig. 36). “ab. nigra Frank” //“Nordl. Kalkalpen / Chiemgau / Lofer/ 28.V.1920 / coll. Dr. Gelpke” // “Type” // “ab. nigra Frank” // “Mus. Altona comm. / Eing Nr 3 - 65” // “ZMH 824709”.
Original locality. Nordl. Kalkalpen [Austria].
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hart- manni Standfuss, 1888.
37. ab. punctata Frank
Original combination. Parnassius mnemosyne hart- manni ab. punctata Frank.
Evolutionary Systematics 5 2021, 193-261 207
i Slg.Mus.Altona Eing.Nr.6/1978 ZMH 824715
Slg-Muse Altona Eing.Nr.6/1978
ZMH 824716
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2MH 824712
Plate 6. 31. Parnassius epaphus beicki Bryk & Eisner, 1932; 32. Parnassius epaphus ab. rubropicta Bang-Haas, 1915; 33. Parnas- sius delphius candidatus Bang-Haas, 1915; 34. Parnassius jacobsoni eva Kotzsch, 1936; 35. Parnassius mnemosyne hartmanni ab. minor Frank; 36. Parnassius mnemosyne hartmanni ab. nigra Frank; 37. Parnassius mnemosyne hartmanni ab. punctata Frank; 38. Parnassius mnemosyne hartmanni ab. nigerrima Frank. a. Dorsal view, b. ventral view.
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208 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 1’, “Co- type” 14 (ZMH 824710-824711) (Fig. 37). “Cotypen” // “Nordl. Kalkalpen / Inntaler Berge / Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // “Mus. Alto- na comm. / Eing. Nr. 3-65” // “ZMH 824710”; “Typen” // “Mus. Altona comm. / Eing. Nr. 3-65” // “Nordl. Kalkalpen / Inntaler Berge / Oberaudorf / 30.V.23 500- 700 m / coll. Dr. Gelpke” // “ab. punctate Frank” // “ZMH 824711”.
Original locality. Nordl. Kalkalpen [Austria].
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hart- manni Standfuss, 1888.
38. ab. nigerrima Frank
Original combination. Parnassius mnemosyne hart- manni ab. nigerrima Frank.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 14 (ZMH 824712) (Fig. 38). “Nordl. Kalkalpen / Inntaler Berge / Oberaudorf / 1.VI.24 500-700 m / coll. Dr. Gelpke” // “Type” // “Mus. Altona comm. / Eing. Nr. 3-65” // “ab. nigerrima Frank” // “ZMH 824712”.
Original locality. Nordl. Kalkalpen [Austria].
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable. The current name of the subspecies is Parnassius mnemosyne hart- manni Standfuss, 1888.
39. ab. subtusalbipupillata
Original combination. Parnassius apollo ab. subtusal- bipupillata.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 1¢ (ZMH 824720) (Fig. 39). “Balkan / Bulgaria / Sofia / Mt. Vi- tosch / 2000 m Jul” // “ab. quincunx Bryk / ab. reniformis O. B.-H. / ab. laticincta Trti.” // “Mus. Altona comm. / Eing. Nr. 3-65” // “Type” // ZMH 824720”.
Original locality. Balkan, Bulgaria.
Remarks. This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable. The current name of the species 1s Parnassius apollo (Linnaeus, 1758).
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40. ab. analifasciata Gelpke
Original combination. Parnassius apollo ab. analifas- ciata Gelpke.
Current status. “in litteris’ name and hence not available.
Original material. Labelled as “Type” 1¢ (ZMH 824722) (Fig. 40). “Stiddeutschland / Frank. Jura / Eich- statt / 12.VII.1921 / coll. Dr. Gelpke” // “Type” // “ab. analifasciata Gelpke” // “ab. splendida Osth. —fasciata st. / ab. magnifica Bryk / ab. subt-bip. Fr. ab. subt-graphica / ab. seitzi Brk. — embryki Bryk.” // “Mus. Altona comm. / Eing. Nr. 3-65” // “ZMH 824722”.
Original locality. South Germany.
Remarks. This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999); it is deemed to be an infrasubspecific name (the author used “aberra- tion”, “ab.”) and is hence unavailable. The current name of the species is Parnassius apollo (Linnaeus, 1758).
Subfamily Papilioninae Latreille, 1802 Tribe Papilionini Latreille, 1802
41. f. unimaculata Warnecke, 1924
Original combination. “Papilio xuthus Linnaeus f. uni- maculata” Warnecke, 1924 Dt. Ent. Z. Iris 38: 149.
Current combination. Papilio xuthus f. unimacula- ta Warnecke, 1924.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 1¢ (ZMH 61623) (Fig. 41). “Japan” // “Type” // Sammlung / Fritz Diehl / Eing. Nr. 35, 1950” // “P. xuthus L. / n.f. // uni- maculata / Warn.” // “ZMH 61623”.
Original locality. Original publication without loca- tion records; label states Japan; according to hand written type catalogue of Warnecke: Ussuri ex Coll. Dorries (67), holotype destroyed in fires in Zool. Mus. Hamburg 1943, 1 specimen with location label Japan labelled as type by Warnecke in Coll. Warnecke, now in ZMH.
Remarks. Warnecke (1924) proposed this name as a form of P. xuthus Linnaeus, 1776. According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and 1s hence un- available. The text of Warnecke (1942) paper unambigu- ously reveals that he describes an individual form and not a (geographic) subspecies.
Tribe Troidini Talbot, 1939
42. peruanus Fuchs, 1954
Original combination. “Papilio archidamas var. peruanus” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 83-84. Current combination. Battus polydamas peruanus (Fuchs, 1954). Current status. Valid subspecies.
Evolutionary Systematics 5 2021, 193-261
Type material. Lectotype 14 (ZMH 61615) (Fig. 42) Paralectotypes 2312 (ZMH 61616-61617, ZMH 824999). “Rio Pampas / Cuesta de Bonbon / 8/1928 / ca. 2000 m / Type” // “HOLOTYPE / Papilio archidamas / var. peruanus / von Fuchs” // “archidamas / S. Rio Pam- pas / 2000 m” // “Z.1.M. / Hamburg” // “LECTOTYPE 3 / Papilio archidamas / var. peruanus Fuchs / by G. La- mas, 1978” // “Coll. H. Rédinger / Eing. Nr. 30, 1952 / Katalog Nr. L.” // “Battus archidamas / peruanus von Fuchs / G. Lamas. Det. 1978” // “Battus polydamas / pe- ruanus (Fuchs, 1954) / Lectotype 3 / Racheli & Pisched- da 1987” // “ZMH 61615”; “Rio Pampas / Cuesta de Bonbon / 8/1928 / ca. 2000 m / Type” // “P. archidamas peruanus” // “Z.1.M. Hamburg” // “Battus polydamas / peruanus (Fuchs, 1954) / Paralectotype 3 / Racheli & Pischedda, 1987” // “ZMH 61616”; “Rio Pampas / Cues- ta de Bonbon / 8/1928 / ca. 2000 m / Type” // “Battus polydamas | peruanus (Fuchs, 1954) / Paralectotype 9 / Racheli & Pischedda, 1987” // “Z.1.M. Hamburg” // “ZMH 61617”; “3” // “Papilio / archidamas & / var. pe- ruanus / von Fuchs / Rio Pampas / Cuesta de Bonbon / c. 2000 m / 8/28” // “Z.I.M. / Hamburg” // “Type” // “Battus polydamas / peruanus (Fuchs, 1954) / Paralectotype 3 / Racheli & Pischedda 1987” // “Coll. H. Rodinger / Eing. Nr. 30, 1952 / Katalog Nr. L.” // “ZMH 824999”.
Type locality. [Lectotype] Peru: “Rio Pampas, Cues- ta de Bonbon, ca. 2000 m”; lectotype designated by La- mas (1978).
Remarks. Fuchs (1954) proposed this name as a va- riety of P. archidamas Boisduval, 1836. According to article 45.6.4 (ICZN 1999), it is subspecific if first pub- lished before 1961 and author expressly used the terms “variety” or “form”. Papilio archidamas Boisduval, 1836 is a junior subjective synonym of Battus polydamas (Lin- naeus, 1758).
Tribe Leptocircini Kirby, 1896 43. ab. ornatissima Warnecke, 1924
Original combination. “Papilio podalirius L. n. ab. or- natissima Warn.” Warnecke, 1924 Int. Ent. Z. 18: 157.
Current combination. [phiclides podalirius ab. or- natissima (Warnecke, 1924).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 61618) (Fig. 43). “n. ab. ornatissima / Warnecke. Type” // “Type” // “altester [?] Name: confluens Ver. / Rhop. Pal. P. 292, t. 57, f. 5, 1911 / vgl Lambillioneaa 1932 p. 223” // “Katalog Nr. / L. 204” // “Slg. G. Warnecke / eing. Nr. 5. 1949” // “ZMH 61618”.
Original locality. Southern Germany.
Remarks. Warnecke, (1924) proposed this name as an aberration of Papilio podalirius Linnaeus, 1758. There- fore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.
209 Unpublished, unavailable names
44. ab. indistinctus Meinhard
Original combination. [phiclides feisthamelii f. ornat- us ab. indistinctus Meinhard.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 14 (ZMH 835574) (Fig. 44). “26/5. 1906 / Stuttgart / ex larv.” // “ab. indistinctus / Meinh.” // “ZMH 835574”.
Remarks. This taxon cannot be found in the literature. Probably an aberration of Jphiclides feisthamelii (Du- ponchel, 1832). Apparently the imago emerged in Stutt- gart (Germany), but the larva must have been collected in the Iberian Peninsula (M. Wiemers pers.com. ).
Family Pieridae Swainson, 1820 Subfamily Coliadinae Swainson, 1827
45. ab. crocopepla Warnecke, 1912
Original combination. “Colias aurora Esp. 9 ab. croco- pepla” Warnecke, 1912 Soc. Ent. 27: 68.
Current combination. Colias heos ab. crocopepla Warnecke, 1912.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type’12 (ZMH 827594) (Fig. 45). “Type” // “Bikienfl.” // “ab. crocope- pla / Warn. Soc. Ent. 27 6. 68” // “L.6” // “sig. G. War- necke / Eing. Nr. 5, 1949” // “Katalog Nr. / L 1754” // “AMUR-GEBIET: / Gebr. Dorries leg. 1887 / Erworb. e. coll. / Doérries sen. 1917 / (vgl. Mém. Rom. VI, / 1892 p 91)” // “ZMH827594”.
Original locality. Amur [Russia].
Remarks. Papilio aurora Esper, 1783 is a junior pri- mary homonym of Papilio aurora Cramer, [1780]. The former is currently known as Colias heos (Herbst, 1792) and the latter is known as Colotis aurora (Cramer, [1780]). Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias aurora [ab.| croco- pepla is infrasubspecific and thus not available name.
46. ab. rhododactyla Warnecke, 1912
Original combination. “Colias aurora Esp. 9 ab. rhodo- dactyla” Warnecke, 1912 Soc. Ent. 27: 68.
Current combination. Colias heos ab. rhododactyla Warnecke, 1912.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 192 (ZMH 827593) (Fig. 46). “L.39 / Col. aurora/ ab. 2 rhododactyla / Warn.” // “ab. rhododactyla / Warnecke / Soc. Ent. 27, 1912 p. 68” //“Katalog Nr. /L. 1755” //“Tjutyuché / Ost-Siberien” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827593”.
Original locality. Amur [Russia].
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210 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Remarks. Papilio aurora Esper, 1783 is a junior pri- mary homonym of Papilio aurora Cramer, [1780]. The former is currently known as Colias heos (Herbst, 1792) and the latter is known as Colotis aurora (Cramer, [1780]). Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias aurora [ab.| rhodo- dactyla is infrasubspecific and thus not available name.
47. ab. theia Warnecke, 1912
Original combination. “Colias aurora Esp. & ab. theia” Warnecke, 1912 Soc. Ent. 27: 68.
Current combination. Colias heos ab. theia War- necke, 1912.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Lectotype” 12° (ZMH 827609) (Fig. 47). “Kentei- / Gebirge” // “Gebr. Dorries leg.1889 / Erworb. E. coll. / Dorries sen. 1917” // “f. theia Warnecke / Die Type ist im Zool. Mus. Hamburg!” // “L. 38 / Lekto-Type / Col. aurora f. theia warn.” // “Katalog Nr. /L 1750” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827609”.
Original locality. Amur [Russia].
Remarks. Papilio aurora Esper, 1783 is a junior pri- mary homonym of Papilio aurora Cramer, [1780]. The former is currently known as Colias heos (Herbst, 1792) and the latter is known as Colotis aurora (Cramer, [1780]). Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stated that Colias aurora [ab.] theia is infrasubspecific and thus not an available name.
48. f. rebeli Schawerda, 1906
Original combination. “Colias myrmidone balcdnica f. rebeli” Schawerda, 1906; Verh. Zool. Bot. Ges. Wien 6: 651.
Current combination. Colias caucasica balcanica f. rebeli Schawerda, 1906.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 12 (ZMH 827619) (Fig. 48). “Orjen / Herzegowina / Dr. Schaw- erda” // “Sammlung / Fritz Diehl / Eing. Nr. 35,1950” // “rebeli 2” // “Cotype” // “Cotype” // “LS” // “Katalog Nr. L 1721” // “ZMH 827619”.
Original locality. Bosnia and Herzegovina: Trebevic”’.
Remarks. Schawerda (1906) proposed this name as a form of C. m. balcanica Rebel, 1901. Therefore, as stat- ed by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infra- subspecific rank) and 1s hence unavailable. Colias bal- canica Rebel, 1901 is currently regarded as subspecies
(<4
vom
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of C. caucasica Staudinger, 1871. Grieshuber and Lamas (2007) provided a nomenclatural list of all available and unavailable names in the genus Colias and clearly stat- ed that Colias myrmidone balcanica |f.| rebeli is infra- subspecific and thus not available name.
49. tatarica Bang-Haas, 1915
Original combination. “Colias cocandica tatarica O. B.- H. n. var.” Bang-Haas, 1915 Dt. Ent. Z. Iris 29: 98.
Current combination. Colias cocandica tatarica Bang-Haas, 1915.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 833446) (Fig. 49). “Co-Type” // “Chotan mer / Karakorum / Juli” // “VII. / Chotan. mer. / Karakorum / v. Friedrich [?]” // “ZMH 833446”.
Type locality. China: “... am Nordabhange des Kara- korum, in den Gebirgen stidlich der Stadt Chotan (Chi- nesische Tatarei)” [...around the northern slopes of the Karakorum, in the Mountains south of the town of Cho- tan (Chinese Tartare) ].
Remarks. Bang-Haas (1915) proposed this name as a variety of C. cocandica Erschoff, 1874. According to article 45.6.4 (ICZN 1999), it is subspecific if first pub- lished before 1961 and author expressly used the terms “variety” or “form”.
50. aurea Kotzsch, 1936
Original combination. “Colias wiskotti aurea subsp. n.” Kotzsch, 1936 Ent. Rdsch. 54(4): 45.
Current combination. Colias wiskotti aurea Kot- zsch, 1936.
Current status. Valid subspecies.
Type material. Syntypes 3¢¢ 19 (ZMH 827622- 827623, ZMH 835580—-835581) (Fig. 50). “Nord-Ost- Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte Juli/leg. H. & E. Kotzsch” // “Wisk. aurea Kotzsch” // “Co-Type / e Collection / Kotzsch” // “L.7” //“Katalog Nr. /L 1793” //“Sig G. Warnecke / Eing. Nr. 5, 1946” // “ZMH 827622”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500—4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Katalog Nr. / L 1794” // “Wiskotti aurea Ktz” // “Co-Type / e Collection / Kotzsch” //“L.8” // “Sig G. Warnecke / Eing. Nr. 5, 1946” // “ZMH 827623”; “Nord-Ost-Hindukusch / Nuksan- Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Co-Type / e Collection / Kotzsch” // “Wiskotti 4/ aurea’ // “ZMH 835580”: “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H. & E. Kotzsch” // “Colias 9° / wiskotti / v. aurea” // “Co-Type / e Collection / Kotzsch” // “ZMH 835581”.
Type locality. Afghanistan:“Ost-Hindukusch, Nord- seite, Alpenwiesenzone, 3500-3800 m.”
Evolutionary Systematics 5 2021, 193-261 211
Siiddeutschland Frank. Jura he
Eichstatt v7
Balkan: coll. Dr, Gelpke sgzy BULGARIA et | — 2000 aul | | Pp yD e | | ab. quincunx rac 4b. seniors OB.
| abanalifasciala Gelnas
a6. spon dicta dst fasctala. af sabtbsn takes repli ab sectry Bat emBryne be
Mus. A tona comm | ting. NF, 3-64 ZMH 824722
ab laticinclareh.
Mus. Altona comin Fing Ne 3+ 65
Type _ |
[zm 824720
ee Z.bM. Hamner |
4
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| Battus pelydseas | pervanvs (Fecks 1454) Lectotypéee? Rachel’ & Pischeclela (98%
ry
ZMH * 61615
|ZMH 835574
Plate 7. 39. Parnassius apollo ab. subtusalbipupillata; 40. Parnassius apollo ab. analifasciata Gelpke; 41. Papilio xuthus f. unimac- ulata Warnecke, 1924; 42. Battus polydamas peruanus (Fuchs, 1954); 43. Iphiclides podalirius ab. ornatissima (Warnecke, 1924); 44. Iphiclides feisthamelii f. ornatus ab. indistinctus Meinhard. a. Dorsal view, b. ventral view.
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Z2N2 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
ZMH 827593
Orjen 3. Herzegowina
“ Dr.Sghawerda
Sammlung : Eritz Diehl + Eing.Nr95,1950 !
ZMH 827609
ZMH 827619
Nord-Ost-Hindukuseh_ Nuksan-Pass-Nordseite : enwitsenzone
: Mitte Juli leg.H.&EKotzsch
sult bs
11cm
Plate 8. 45. Colias heos ab. crocopepla Warnecke, 1912; 46. Colias heos ab. rhododactyla Warnecke, 1912; 47. Colias heos ab. theia Warnecke, 1912; 48. Colias caucasica balcanica f. rebeli Schawerda, 1906; 49. Colias cocandica tatarica Bang-Haas, 1915;
50. Colias wiskotti aurea Kotzsch, 1936. a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
51. yauta Fuchs, 1928
Original combination. “T7eriocolias yauta’ Fuchs, 1954 Beitrage zur Fauna Perus 4:83.
Current combination. Eurema Zelia andina (Fuchs, 1928).
Current status. Junior subjective synonym of Eurema zelia andina (Forbes, 1928).
Type material. Holotype 14 (ZMH 825001), Para- types 80:2 (ZMH 825000, ZMH 825002-825008) (Fig. 51). “Teriocolias / yauta 3 / J. Fuchs / Cordillere von / Tlaula 5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “Type” // “Paratype / Teriocolias yauta / von Fuchs” // “Teriocolias zelia | andina Forbes / G. La- mas. det. 1978” // “ZMH 825000”; “Llauta / ca. 3000 m / 5/1922 / Type” // “Coll. H. R6dinger / Eing. Nr. 30, 1952” // “Holotype / Teriocolias yauta / von Fuchs” // “Terioco- lias zelia | andina Forbes / G. Lamas. det. 1978” // “ZMH 825001”; “Llatita / ca. 3000 m / 5/1922 / Type” // “Coll. H. Rodinger / Eing. Nr. 30, 1952” // “ZMH 825002”; “Lara- mate / ca 3400 m / 5/1922 / Type” // “Coll. H. Roding- er / Eing. Nr. 30, 1952” // “ZMH 825003”; “Oberhalb / Aina / ca 3000 m / 8/1924 / Type” // “Coll. H. R6dinger / Eing. Nr. 30, 1952” // “ZMH 825004”; “Llatita / ca. 3000 m / 5/1922 / Type” // “Coll. H. Rodinger / Eing. Nr. 30, 1952” // “ZMH 825005”; “Type” // Teriocolias / yauta 9 / v. Fuchs / Cordillere von / Ilaula 5.22 / 3000 m” // “Sam- mlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “ZMH 825006”; “Type” // “Sid — Peru / Ocana, 2600 m / 7.4.1936.” // “Hbg. — Std — Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” //“Teriocolias / yauta 3 / v. Fuchs” // “Sammlung / W, v Fuchs / Eing. Nr. 4, 1956” // “ZMH 825007”; “Type” /! Teriocolias | yauta 2 / v. Fuchs / Cordillere von / Taula 5.22 / 3000 m” // “Sammlung / W.v. Fuchs / Eing. Nr. 4. 1956” // “ZMH 825008”.
Type locality. South Peru. “ ... Kordillere von Llauta ... auch an anderen Stellen der Kordilleren” ” [Cordillera of Llauta...also other areas in the cordilleras].
Subfamily Dismorphiinae Schatz, 1887
52. lorkovici Pfeiffer, 1932
Original combination. “Leptidea duponcheli Stgr. ssp. n. Jorkovici m.” Pfeiffer, 1932 in Osthelder and Pfeiffer 1932, Mitt. Munch. Ent. Ges. 22: 20.
Current combination. Leptidea duponcheli lorkovici Pfeiffer, 1932.
Current status. Valid subspecies.
Type material. Syntypes 244 (ZMH 827614827615) (Fig. 52). “Syr. Sept. / Taurus c. / Marasch / 6-900 m V.30 / Einh. Slr. leg.” // “Cotypus / duponcheli / ssp. lorkovici / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827614”; “Syria s. / Amanus s. / (Duldtil Dagh) / Jeschildere / VI 33 / Coll. Pfeiffer / Mtinchen” // “Coty- pus / duponcheli / ssp. lorkovici / Pfeiffer” // “Coll. By- tinski-Salz / Eing. Nr 20 - 60” // “ZMH 827615”.
Type locality. Turkey: “Marasch in tirkisch Nordsy- rien; An den Héngen um M bis 1200 m” [Marasch in the
213
turkish regions of northern Syria. On the slopes around Marasch up to 1200 m].
53. f. fragilis Dannehl, 1925
Original combination. “Leptidea sinapis L. (gen. autum- nalis) v. fragilis Dannehl” Dannehl, 1925 Ent. Z. 39: 6.
Current combination. Leptidea sinapis f. fragilis Dannehl, 1925.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 14 (ZMH 835575) (Fig. 53). “3.10.24” // “Sudtirol Terlan / Franz Dannehl” // “fragilis Dhl. ” // “Cotype” // “3.10 1924 / Terlan / Siidtirol / Dannehl” // “ZMH 835575”.
Original locality. Italy: “in Stdtirol bei Bozen, Fle- imstal, Nonstal.”
Remarks. Dannehl (1925) proposed this name as a form of L. sinapis (Linnaeus, 1758). According to article 45.6.1 ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was pro- posed for an infrasubspecific entity, and is hence unavail- able. The text of Dannehl (1925) paper unambiguously reveals that this form was described as a seasonal form (autumn generation) and not as a subspecies.
54. ab. simbruina Dannehl, 1927
Original combination. “Leptidea sinapis L. ab. simbru- ina Dhl.” Dannehl, 1927 Mitt. Mtinch. Ent. Ges. 17: 1.
Current combination. Leptidea sinapis ab. simbru- ina Dannehl, 1927.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 2¢¢ (ZMH 835582—835583) (Fig. 54). “25.6” // “Italia central. / Mti. Simbruini / coll. F. Dannehl” // “simbruina” // “Cotypen” /1 “25.6. 1927 / Mte Simbruini / Central Italien” // “ZMH 835582”; “7.4. 1927 / Mte Simbruini / Central Italien” // “ZMH 835583”.
Original locality. Italy: “Sabiner und Italy: Simbru- iner Berge vom Tal (Campagna) bis auf die Gipfel der angegebenen Gebirge.” [Sabiner and Italy: Simbruin Mountains from the valley (Campagna) up to the top of the stated mountain ranges].
Remarks. Dannehl (1927) proposed this name as an aberration of L. sinapis (Linnaeus, 1758). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used one of the terms “aberration”, “ab.” or “morph”) and is hence unavailable.
55. stabiarum Staudinger, 1914
Original combination. “Leptidea (Leucophasia) sina- pis stabiarum m., nov. subspec.” Staudinger, 1914 Zeit. Wiss. Ins. Biol. 10: 371, t. 11, figs. 5, 6.
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Current combination. Leptidea sinapis stabiarum Staudinger, 1914.
Current status. Junior subjective synonym of nomi- notypical Leptidea sinapis (Linnaeus, 1758).
Type material. Syntypes 3¢¢ (ZMH 827624— 827626) (Fig. 55). “Ital. mer. 600 m. / penins. Surren- tina / M. Faito 3.6.1920 / H. Stauder legit.” // “Type” // “Sammlung / G Warnecke / Eing Nr 5, 1949” // “ZMH 827624”; “25/6 / Ital. mer. m. / penins. Surrentina / M. Fatio 3.6.1920 / H. Stauder legit.” // “Type” // “Sammlu- ng / G Warnecke / Eing Nr 5, 1949” // “ZMH 827625”; 26/6 / Ital. mer. 700 m. / penins. Surrentina / M. Fatio 3.6.1920 / H. Stauder legit.” // “Type” // “Sinapis stabi- arum / Stauder / v. stabiarum.” // “ZMH 827626”.
Type locality. Italy: Monte Faito and S. Angelo [Na- ples], 700-1400 m.
Subfamily Pierinae Swainson, 1820 56. italorum Dannehl, 1929
Original combination. “Anthocharis euphenoides St. italo- rum Dhl.” Dannehl, 1929 Mitt. Munch. Ent. Ges. 19: 98.
Current combination. Anthocharis euphenoides italorum Dannehl, 1929.
Current status. Junior subjective synonym of nominotypical Anthocharis euphenoides Staudinger, 1869.
Type material. Syntype 14 (ZMH 835584) (Fig. 56). “20.5” // “Apenn. Central. / Mte. Genzana / 1500—2000 m. / coll. F. Dannehl” // “Cotype” // “20.5.1928 / Mte Genzana / Central- Apenninen / 1500-2000 m D.” // “ZMH 835584”.
Type locality. Italy: in den Gebieten des Velino, Sirente, Monte Genzana, Monte Paradiso und der Majella. auf Hohen von 1500-2000 m.” [in the mountain ranges of the Velino, Sirente, Monte Genzana, Monte Paradiso and Majella. In altitudes of 1500-2000 m].
(<4
57. paravicinii Stauder, 1915
Original combination. “Euchloé belia paravicinii”’ Stauder, 1915 Dt. Ent. Z. Iris 29: 25.
Current combination. Euchloe ausonia paravicinii Stauder, 1915.
Current status. Junior subjective synonym of Eu- chloe ausonia melanochloros Rober, 1907.
Type material. Syntype 1¢ (ZMH 827616) (Fig. 57). “Algeria mer. / Djebel Aures / 10.5.1914 / H. Stauder” // “Paravicinii Stauder” // “paravicinii Stauder” // “Type” // “L. 4” // “Katalog Nr. / L. 1180” // “Type” // “Sig. G. Warnecke / Eing. Nr. 5, 1948” // “ZMH 827616”.
Type locality. Algeria: “Djebel Aureés in Stidalgerien”’.
Remarks. Papilio belia Stoll [1782] is a primary hom- onym of Papilio belia Linnaeus, 1767 which is currently known as Anthocharis belia.
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
58. f. flavide apicata Stauder, 1922
Original combination. “7eracolus daira nouna auresia- ca g. II. interposita Stdr. f. flavide apicata m. nov.” Staud- er, 1922 Mitt. Mitinch. Ent. Ges. 12: 24—25.
Current combination. Colotis evagore nouna f. fla- vide apicata (Stauder, 19272).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 74° (ZMH 827595-827599, ZMH 827601-827602) (Fig. 58). “Type” // “El Kautara / auf VI” // “Auresica gen. II. / in- terposita Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827595”: “2” // “Type” // “Algeria mer. / Djebel Aures / VI. 1913 / H. Stauder” // “f. inter- posita | 5.VY’ // “VI form / 3 200” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827597”; “Type” // “El Kautara / auf VI’ // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827596”; “Type” // “El Kau- tara / auf VI” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827598”; “3” // “Type” // “Algeria mer. / Djebel Aures / VI. 1912 / H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827599”; “Type” //“X Dyebel Aures / VII. 1912 H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827601”; “Q” // “Type” // “Algeria mer. / Djebel Aures / VI. 1913 / H. Stauder” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827602”.
Original locality. Not indicated. Probably “Algeria mer. / Djebel Aures”’.
Remarks. Stauder (1922) proposed this name as a form of 7! d. nouna auresiaca interposita Staud- er, 1922. Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and 1s hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).
59. f. vernalis alticola auresiaca Stauder, 1913
Original combination. “T7eracolus daira (= dalia Fldr.) nouna Luc. 1848 (= demagore Fldr.) forma vernalis alti- cola auresiaca” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.
Current combination. Colotis evagore nouna f. ver- nalis alticola auresiaca (Stauder, 1913).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 3939 (ZMH 827600, ZMH 827603-827607) (Fig. 59). “3” // “Co- type” // “Algeria mer. / Djebel Aures / V. 1912 / H. Staud- er” // “auresiaca” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827600”; “2” // “Type” // “Algeria mer. / Djebel Aures / V. 1912 / H. Stauder” // “auresiaca / S Stdr.” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” /1 “ZMH 827603”; “9” // “Type” // “Algeria mer. / Djebel Aures / V. 1912 / H. Stauder” // “auresiaca Stdr. / 2” //
Evolutionary Systematics 5 2021, 193-261 215
Coll. H. ‘Rbdinger Eing. Nr, 30, 1992
TYPE FO otaS gousbe ‘Goll, Bytinski-Salz: you Fucks teing. Ns. 20, 1960: Teyocokkas 2eka Anokne. forbes
G. Lamas. det. 1978 ;
ZMH 825001|
Apenn.central. Mite. Génzana’ 1500-2000 m
hee
nna se
[2M 827616
1 cm
Plate 9. 51. Eurema zelia andina (Fuchs, 1928); 52. Leptidea duponcheli lorkovici Pfeiffer, 1932; 53. Leptidea sinapis f. fragilis Dannehl, 1925; 54. Leptidea sinapis ab. simbruina Dannehl, 1927; 55. Leptidea sinapis stabiarum Staudinger, 1914; 56. Antho- charis euphenoides italorum Dannehl, 1929; 57. Euchloe ausonia paravicinii Stauder, 1915; 58. Colotis evagore nouna f. flavide apicata (Stauder, 1922). a. Dorsal view, b. ventral view.
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“Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827604”: “9” // “Cotype” // “Algeria mer. / Djebel Aures /V. 1912 /H. Stauder” // “auresiaca / 9” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827605”; “2” // “Cotype” // “Algeria mer. / Djebel Aures / V. 1912 / H. Stauder” // “auresiaca / 2” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827605”; “4” // “Type” // “Algeria mer. / Djebel Aures / V. 1912 / H. Stauder” // “auresiaca Stdr. / 3” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827607”.
Original locality. Algeria: “in den algerischen Atlas und die nordliche Sahara” [in the Algerian Atlas and the northern Sahara].
Remarks. Stauder (1913) proposed this name as a form of 7: d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 ICZN 1999) it is deemed to be an in- frasubspecific name (the author expressly gave it infra- subspecific rank) and is hence unavailable. Zeracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).
60. f. biformata Stauder, 1913
Original combination. “T7eracolus daira (= dalia Fldr.) nouna Luc. 1848 (= demagore Fldr.) forma autumnalis biformata m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.
Current combination. Colotis evagore nouna f. bi- formata (Stauder, 1913).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827608) (Fig. 60). “4” // “biformata / < Stdr.” // “Type” // “Algeria mer. / Djebel Aures / V. 1912 /H. Stauder” //“Sam- mlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827608”.
Original locality. Algeria: “in den algerischen Atlas und die nordliche Sahara” [in the Algerian Atlas and the northern Sahara].
Remarks. Stauder (1913) proposed this name as a form of 7! d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an in- frasubspecific name (the author expressly gave it infra- subspecific rank) and is hence unavailable. Teracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).
61. f. aestiva pyroleuca Stauder, 1913
Original combination. “7eracolus daira nouna Luc. for- ma aestiva pyroleuca m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.
Current combination. Colotis evagore nouna f. aes- tiva pyroleuca (Stauder, 1913).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type and Cotype” 1049 (ZMH 827618, ZMH 827628-827636) (Fig. 61).
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
“O” // “Dyjebel Aures / VII. 1912 Stauder” // “nouna var. / pyroleuca / Stauder” // “TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827618”; “2” // “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // “pyroleuca 2” //“COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827628”; “3” // “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // “Cotype” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827629”, “3” // “Algeria mer. / Djeb- el Aurés / VII. 1912 / H. Stauder” // “pyroleuca 4” // “TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” /1 “ZMH 827630”; “2” // “Algeria mer. / Djebel Aurés / VII. 1912 /H. Stauder” // “pyroleuca 9” // ““COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827631”: “3” // “Algeria mer. / Djebel Aurés / VII. 1912 / H. Stauder” // “pyroleuca 6” // “COTY PE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827632”; “2” // “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // “pyroleuca 3 NO” // “COTYPE” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827633”; “3” // “COTYPE” // Algeria mer. / El Kaictara [?] / anfg [IX 19 / H. Stauder” // “pyroleuca 4” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827634”; “9” // “Type” // “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // “pyroleuca 2” // “Sammlung / G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827635”; “3” // “TYPE” // “Algeria mer. / Djebel Aures / VII. 1913 /H. Stauder” // “Sammlung /G Warnecke / Eing. Nr. 5, 1949” // “ZMH 827636”.
Original locality. Algeria: “in den algerischen Atlas und die nordliche Sahara” [in the Algerian Atlas and the northern Sahara].
Remarks. Stauder (1913) proposed this name as a form of 7! d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (CZN 1999) it is deemed to be an in- frasubspecific name (the author expressly gave it infra- subspecific rank) and is hence unavailable. 7eracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).
62. ab. evagorides Stauder, 1913
Original combination. “Zeracolus daira nouna Luc. ab. evagorides m.” Stauder, 1913 Zeit. Wiss. Ins. Biol. 9: 290.
Current combination. Colotis evagore nouna ab. evagorides (Stauder, 1913).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 2¢.¢ (ZMH 827620-827621) (Fig. 62). “4” // “Algeria mer. / Djebel Aureés / VI. 1913/H. Stauder” // “Sammlung /G Warnecke / Eing. Nr. 5,1949” // “COTYPE” // “ZMH 827620”; “3” // “evagorides / Stdr.” // “Algeria mer. / Djebel Aures / VII. 1912 / H. Stauder” // “TYPE” // “Sammlung / G Warnecke / Eing. Nr. 5,1949” // “ZMH 827621”.
Original locality. Algeria: “in den algerischen Atlas und die nordliche Sahara” [in the Algerian Atlas and the northern Sahara].
Evolutionary Systematics 5 2021, 193-261
Remarks. Stauder (1913) proposed this name as an aberation of 7’ d. nouna (Lucas, 1848). Therefore, as stated by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. 7eracolus daira nouna Lucas, 1849 is junior subjective synonym of Colotis evagore nouna (Lucas, 1849).
63. italorum Stauder, 1921
Original combination. “Pieris brassicae L. f. n. (g. aest. mer. alticola) italorum” Stauder, 1921 Dt. Ent. Z. Iris 35: 26.
Current combination. Pieris Stauder, 1921.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH 827617) (Fig. 63). “Calabria mer. / Aspromonte 900 m. / 29/6 Polsi [?] 1920 /H. Stauder Legit.” // “Type” // “brassicae var. / italorum Staud.” // “ZMH 827617”.
Type locality. Italy: “1.[talien] c.[entralis] m.[eridiona- lis], Aspromonte, Montalto, 1800 m.”
brassicae italorum
64. elbursina Bytinski-Salz & Brandt, 1937
Original combination. “Pieris ergane, Hbn. ssp. e/bursina, ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: 1-2.
Current combination. Pieris ergane elbursina By- tinski-Salz & Brandt, 1937.
Current status. Valid subspecies.
Type material. Syntypes 34'3' 19 (ZMH 827610- 827613) (Fig. 64). “IRAN / Keredj (1500 m.) / 18/5 1936 / Brandt” // “Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827610”; “TRAN / Keredj (1200 m.)/ 4.4. 1936 / Brandt” // “Cotypus / P. ergane / ssp. pfeifferi / By.S & Br.” // “ssp. pfeifferi Byt.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827611”; “IRAN / Keredj (1500 m.) / 20/5 1936 / Brandt” //“Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “Coll By- tinski-Salz / Eing. Nr. 20, 1960” // ZMH 827612”; “IRAN / Keredj (1500 m.)/ 18/5 1936 / Brandt” // “Cotypus / ergane ssp. / pfeifferi / By.S+Br.” // “ssp. pfeifferi / By.S. & Br.” // “Coll Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827613”.
Type locality. Iran: Tehran, “Keredj” [Karaj].
Remarks. Label indicates ssp. pfeifferi, but apparent- ly this name does not exist. Collection data are identical to ssp. e/bursina, possibly a last-minute name change by Bytinski-Salz.
65. r. exigua Verity, 1923
Original combination. “Pieris ergane, H.-G. race exi- gua, mihi.” Verity, 1923 Ent. Rec. 35 Suppl. (18).
Current combination. Pieris ergane r. exigua Veri- ty, 1923.
217
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Cotype” 244 (ZMH 835577—-835578) (Fig. 65). “exigua Vrty” // “Marche / Bolognola / 7 Ag 1937 / Querci” // [blank label] // “ZMH 835577”; “exigua Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 835578”.
Original locality. Italy: “Upper Fargno Valley, at 1400 m., in the Sibillini Mts.”
Remarks. Verity (1923) proposed this name as a race of P. ergane (Geyer, 1828). According to Kudrna (1983), this name in unavailable because it is a seasonal form and also a nomen nudum. The year on the specimen labels (1937) indicate specimens were collected 14 years after description (1923) from the same location the original holotype material was collected. As a result, the black la- bel of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called to- potype. According to ICZN (1999), a topotype has no for- mal standing and is not regulated by the Code. Therefore, these two specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
66. f. rossioides Stauder, 1921
Original combination. “P. rapae L. f. n. rossioides” Stauder, 1921 Dt. Ent. Z. Iris 35: 27.
Current combination. Pieris rapae f. rossioides Stauder, 1921.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827627) (Fig. 66). “Calabria mer. / Aspromonte 800 m. / 4/7 [illegible] 1920 / H. Stauder legit.” // “Type” // “Rossioides Stdr.” // “rossioides / Stdr. / Type” // “ZMH 827627”.
Original locality. Italy: “Polsibeken, Aspromonte 7-900 m.”
Remarks. Stauder (1921) proposed this name as a form of P. rapae (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work (please see below) unambiguously reveals that the name was proposed for an infrasubspecific en- tity. In the same paper, Stauder (1921) described another “form” of P. rapae as “zelleri” and clearly stated that it also occurs at the same “type locality” as rossioides (1.e., Aspromonte). As by definition two geographical races (or forms) cannot occur sympatrically; therefore, the content of the work reveals that such forms were not meant to be subspecific. However, Stauder added further complexity; there is a difference between how “zelleri’” and “rossioi- des” have been described. In fact, the former was as ‘g. aest. mer. alt.’, 1.e., generatio aestivalis meridionalis alti- tudinalis [Southern altitudinal summer generation], while the latter just as ‘f. n.’, 1.e., “forma nova” [new form].
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218 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Pn te : fas 61a Gaae | Sue
ZMH 827618|
7
hee ee
‘Coli ee Eimg_ Nr. 20, 1960
ZMH_ 835577
1cm
Plate 10. 59. Colotis evagore nouna f. vernalis alticola auresiaca (Stauder, 1913); 60. Colotis evagore nouna f. biformata (Stauder, 1913); 61. Colotis evagore nouna f. aestiva pyroleuca (Stauder, 1913); 62. Colotis evagore nouna ab. evagorides (Stauder, 1913); 63. Pieris brassicae italorum Stauder, 1921; 64. Pieris ergane elbursina Bytinski-Salz & Brandt, 1937; 65. Pieris ergane r. exigua Verity, 1923. a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
Thus, it could theoretically be argued that rossioides was meant as a geographical race, while zelleri would be a seasonal variation of that. However, this speculation does not seem to be correct, because it makes no sense that someone described a seasonal form of a subspecies be- fore describing the subspecies itself. As a result, looking at the paper as a whole and not to the isolated description of rossioides alone, this name has to be considered as in- frasubspecific (Alberto Zilli pers. comm. ).
Unpublished, unavailable names
67. ab. gottingensis Meinhard
Original combination. Pieris napi v. napaeae ab. got- tingensis Meinhard.
Current status. “in litteris’ name and hence not available.
Original material. Labelled as “Type” 14 (ZMH 835579) (Fig. 67). “v. gottingensis / Mht” // “8/5 1909 / Gottingen / Hainholz” // “ZMH 835579”.
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable.
68. ab. nigromarginata Meinhard
Original combination. Pieris brassicae ab. nigromar- ginata Meinhard.
Current status. “in litteris’ name and hence not available.
Original material. Labelled as “Type” 14 (ZMH 835576) (Fig. 68). “13/8 1916 / Gottingen / Hainberg” // “ZMH 835576”.
Remarks. This taxon cannot be found in the literature. As Stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and 1s hence unavailable.
Family Lycaenidae Leach, 1815 Subfamily Aphnaeinae Distant, 1884
69. dueldueli Pfeiffer, 1932
Original combination. “Cigaritis acamas Klug. ssp. n. dildiili m.” Pfeiffer, in Osthelder and Pfeiffer 1932, Mitt. Munch. Ent. Ges. 22: 38.
Current combination. Cigaritis acamas dueldueli Pfeiffer, 1932.
Current status. Valid subspecies.
Type material. Syntypes 3¢¢ (ZMH 827553- 827555) (Fig. 69). “Syria s. / amanus / Duldtl Dagh / VII.30” // “Cotypus / A. epargyros / ssp. diildiili / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // ZMH 827553”; “Syria sept. / amanus sept. / Duldiul Dagh / VII.30” // “Cotypus / A. epargyros / ssp. diildiili
219
/ Pfeiffer” // “Coll. Bytinski Salz / Eing. Nr. 20, 1960” /| “ZMH 827554”; “Syria sept. / amanus sept. / Duldtl Dagh / VH.30” // “Cotypus / A. epargyros / ssp. diildiili / Pfeiffer” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZLMEP8273599" -
Type locality. Turkey: “Marasch in tirkisch Nordsy- rien; Im DD [Dildtl Dagh].”
Subfamily Lycaeninae Leach, 1815 Tribe Lycaenini Leach, 1815
70. f. emarginata Warnecke, 1942
Original combination. “Chrysophanus alciphron Rott., J, n. f emarginata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.
Current combination. Lycaena alciphron f. emargi- nata (Warnecke, 1942).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827527) (Fig. 70). “Schlesw. — Holst. / Kr. / Bockhorn / W. Wolf leg. / 14.7.40” // “Katalog Nr. / L 2322” //“¢ nf. emarginata / Warn. Type / Iris 1942, 103” // “Chrysopha- nus | alciphron / emarginata Warn. / Type / L.31” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827527”.
Originallocality. Germany: “Segeberg’”; Schleswig-Hol- stein publication says Segeberg; the label of the specimen treated as type by Warnecke states Lauenburg crossed out and replaced by Bockhorn.
Remarks. Warnecke (1942) proposed this name as a form of C. alciphron (Rottemburg, 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. Already the title of Warnecke (1942) paper reveals that this form was described as an aberration and not as a subspecies.
71. ab. alba Schoenfeld, 1924
Original combination. “Chrysophanus virgaureae L.-9, alba m., nov. ab.” Schoenfeld, 1924 Int. Ent. Z. 18: 40.
Current combination. Lycaena virgaureae ab. alba (Schoenfeld, 1924).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 12 (ZMH 827586) (Fig. 71). “Bayern / Mitterteich / 24. VHI.1923 / Schonfeld” // [blank label] // “Type / Chr. virgaureae L. / ab. alba Schonfeld /TI. E. Z. Guben 18. Jahrg / 1924. Nr 6. P. 40 / [illegible]” // “ZMH 827586”.
Original locality. Germany: Bavaria.
Remarks. Schoenfeld (1924) proposed this name as an aberration of C. virgaureae (Linnaeus, 1758), there- fore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberra- tion’, “ab.”) and is hence unavailable.
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220 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
72. f. multipunctata Warnecke, 1942
Original combination. “Chrysophanus virgaureae L., 3, n.f. multipunctata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.
Current combination. Lycaena virgaureae f. multi- punctata (Warnecke, 1942).
Current status. Infrasubspecific and hence unavailable.
Original material. Labelled as “Type” 14 (ZMH 827592) (Fig. 72). “n.f. multipunctata / Warn. Type / Iris 1942, 103” //“Chrysoph. virgaureae / multipunctata / Type Warn. / L.30” // “Holstein” // “Katalog Nr. / L. 2196” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827592”.
Original locality. Germany: Holstein.
Remarks. Warnecke (1942) proposed this name as a form of C. virgaureae (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The title and the text of Warnecke (1942) pa- per unambiguously reveals that this form was described as an aberration and not as a subspecies. It was only found in a single male.
73. f. quercii Turati, 1923
Original combination. “Chrysophanus virgaureae emil- ianus f.n. e. quercii f. n.” Turati, 1923 Atti. Soc. Ital. Sci. Nat. 62: 42.
Current combination. Lycaena virgaureae emil- ianus f. quercii (Turati, 1923).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 646 (ZMH 827580-827585) (Fig. 73). “Marche / Bolognola / 15.VI.1937 / Querci” // [blank label] // “quercii Trti.” /1 “ZMH 827580”; “Marche / Bolognola / 15. VII.1937 / Querci” // [blank label] //“quercii Trti.” // “ZMH 827581”; “Marche / Bolognola / 3 Ag 1937 / Querci” // [blank la- bel] // “quercii Trti.” // “ZMH 827582”; “Marche / Bo- lognola / 27. VII.1937 / Querci” // [blank label] // “quer- cil Trti.” // “ZMH 827583”; “Marche / Bolognola / 5 Ag 1937 / Querci” // [blank label] // “quercui Trti.” // “ZMH 827584”; “Marche / Bolognola / 4 Ag 1937 / Querci” // [blank label] // “quercit Trti.” // “ZMH 827585”.
Original locality. Italy: “Monti Sibillini’.
Remarks. Turati (1923) proposed this name as a form of C. v. emilianus Turati, 1923. Therefore, as stat- ed by article 45.6.1 (ICZN 1999) it is deemed to be an infrasubspecific name (the author expressly gave it infra- subspecific rank) and is hence unavailable.
Subfamily Polyommatinae Swainson, 1827 Tribe Polyommatini Swainson, 1827
74. andarabi Forster, 1938
Original combination. “Lycaena pheretiades andarabi ssp. nov.” Forster, 1937 Mitt. Minch. Ent. Ges. 27: 61.
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Current combination. Agriades pheretiades andarabi (Forster, 1938).
Current status. Valid subspecies.
Type material. Paratypes 4¢6 (ZMH 827575- 827578) (Fig. 74). “West-Hindukusch / Andarab / Gerdoll- zone / 4000-4500 m Mitte August / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forst. / W. Forster Munchen” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827575”; “Nord-Ost-Hindukusch / Nuk- san-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster Munchen” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827576”; “West-Hindukusch / Andarab / Gerollzone / 4000-4500 m Mitte August / leg. H.&E. Kotzsch” // “Pa- ra-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster Munchen” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827577”; “West-Hindukusch / Andarab / Geroll- zone / 4000-4500 m Mitte August / leg. H.&E. Kotzsch” // “Para-Type / Lyc. Pheretiades / andarabi Forest. / W. Forster Munchen” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827578”.
Type locality. Afghanistan: “West-Hindukusch, Anda- rab, 4000-4500 m.”
75. f. altera Ziillich, 1929
Original combination. “Lycaena anteros Frr. f. altera” Zullich, 1929 Z. Ost. Ent. Ver. 14: 52.
Current combination. Arvicia anteros f. altera (Ziil- lich, 1929).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type’1¢ (ZMH 827526) (Fig. 75). “BULGARIA OR. / GIOKTEPE 250 M/ MITTE AUG 1928 / COLL. ZUELLICH” // “ExTyp- icalibus / Specimenibus” // “VIHI 1928 / Gidktepe, / Bul- garien” // “ZMH 827526”.
Original locality. Bulgaria: “Rilogebiet, Sidwestbul- garien, 1200 m, und Gioktepe, Stidostbulgarien, 200 m.”
Remarks. Zillich (1929) proposed this name as a form of L. anteros Freyer, 1839. According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and 1s hence un- available. The text of Zullich (1929: 52) unambiguously reveals that this form was described explicitely for the summer generation.
76. r. modicior Verity, 1938
Original combination. “Aricia anteros, Frr. modicior, nom. nov.” Verity, 1938 Ent. Rec. 50(8): Suppl. (5). Current combination. Aricia anteros r. modicior Verity, 1938. Current status. Infrasubspecific and hence unavail- able name.
Evolutionary Systematics 5 2021, 193-261 221
ZMH 835579
ZMH ‘8276271
ociceesy [ZMH 827527|
ZMH -827586 ZMH -827592|
i1cm
Plate 11. 66. Pieris rapae f. rossioides Stauder, 1921; 67. Pieris napi v. napaeae ab. gottingensis Meinhard; 68. Pieris brassicae ab. nigromarginata Meinhard; 69. Cigaritis acamas dueldueli Pfeiffer, 1932; 70. Lycaena alciphron f. emarginata (Warnecke, 1942);
71. Lycaena virgaureae ab. alba (Schoenfeld, 1924); 72. Lycaena virgaureae f. multipunctata (Warnecke, 1942). a. Dorsal view, b. ventral view.
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222
Original material. Labelled as “Type” 233’, (ZMH 827535—-827536) (Fig. 76). “modicior Vrty” // “Macedo- nia / Olympus, 4500’ / July 2, 1935 / Romev” // [blank la- bel] // “ZMH8274535”; “modicior Vrty” // “anteros Frr.” // “Macedonia / Olympus, 4500’ / July14, 1935 / Rome1” // [blank label] // “ZMH8274536”.
Original locality. Greece: “At the highest altitudes of Mt. Olympus, such as at Prionia, 1500 m”.
Remarks. According to Kudrna (1983: 32), this name is unavailable and denotes a race and seasonal form and is hence unavailable.
77. uliginosa Dannehl, 1921
Original combination. “Lycaena argus Schiff. var. uligi- nosa” Dannehl, 1921 Mitt. Minch. Ent. Ges. 11: 39.
Current combination. Plebejus idas uliginosa (Dan- nehl, 1921).
Current status. Junior subjective synonym of Plebe- Jus argus argus (Linnaeus, 1758).
Type material. Syntypes 443, (ZMH 827541- 827542, ZMH 835595-835596) (Fig. 77). “18.7” // “Ob. Bayern / Beuerberg / c. Dannehl 20” // “Argus v. n. / uliginosa” // “Katalog Nr. / L 2562” // “Cotype” // “L.34/Lyc. argus -/v. uliginosa Dannehl / 3 Cotype” // Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827541”: “18.7” // “Ob. Bayern / Beuerberg / c. Dannehl 20” // “vy. uliginosa / Dannehl” // “Cotype” // “Katalog Nr. / L 2561” // “L.35 / Lye. argus - / v. uliginosa Dannehl / 2 Cotype” // Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827542”; “27.6” // Ob. Bayern / Beuerberg / F. Dannehl” // “Cotype” // “27.6. 1921 / Beuerberg / Oberbayern / v. Dannehl” // “ZMH 835595”; “13.7” // Ob. Bayern / Beuerberg / F. Dannehl” // “Cotype” // “13.7. 1921 / Beuerberg / Oberbayern / v. Dannehl” // “LMIEU8 35596".
Type locality. Germany: Bavaria, Beuerberg.
Remarks. Dannehl (1921) proposed this name as a “var.” of L. argus ({[Denis & Schiffermitiller], 1775). L. ar- gus 1S a junior subjective synonym of Plebejus idas Lin- naeus, [1760]. According to article 45.6.4 ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.
78. balcanica Ziillich, 1929
Original combination. “Lycaena argyrognomon Bgstr. f. balcanica” Zillich, 1929 Z. Ost. Ent. Ver. 14: 52.
Current combination. Plebejus argyrognomon bal- canica (Ziillich, 1929).
Current status. Preoccupied by Lycaena semiargus balcanica Tutt, 1909; replacement name Lycaeides ar- gyrognomon baldur (Hemming, 1934) is the current val- id name.
Type material. Holotype 14 (ZMH 827544) (Fig. 78) paratype 146 (ZMH 827543). “BULGARIA
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OCC. // MON. RILSKI 1200 M / 20. — 31 VII. 1928 // COLL. ZUELLICH” // “Ex Typicalibus / Specimini- bus” // “Holotype” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // ZMH 827544”; “BULGARIA OCC. // MON. RILSKI 1400 M / 20 VI. — 6 VII. 1928 // COLL. ZU- ELLICH” // “Ex Typicalibus / Speciminibus” // “Para- typoid” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // ZMH 827543”.
Type locality. Bulgaria: “MON. RILSKI” [Monas- tery Rilski].
Remarks. Zillich (1929) proposed this name as a form of L. argyrognomon (Bergstrasser, 1779). Accord- ing to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “form”.
79. mayencis Eitschberger & Steiniger, 1975
Original combination. “Eumedonia eumedon mayencis n. subsp.” Eitschberger & Steiniger, 1975 Atalanta 6 (1): 106.
Current combination. Eumedonia eumedon may- encis Eitschberger & Steiniger, 1975.
Current status. Valid subspecies.
Type material. Paratypes 546 (ZMH 827548-— 827552) (Fig. 79). “Gonsenheim / 5. Juni / 04 / AN- DREAS / Gonsenheim” // “PARATYPUS 0 / Eumedonia eu- / medon mayencis | EITTSCHBERGER & / STEINI- GER, 1975” //“Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827548”; “Gonsenheim / 5. Juni / 04 / ANDREAS / Gonsenheim” // “PARATYPUS o / Eumedonia eu- / medon mayencis | ETTSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” //“ZMH 827549”; “Mainz / Mombach / 10.6.1922” // “Eumedon” // “PARATYPUS o / Eumedonia eu- / medon mayencis / EITSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827550”; “Gonsenheim / 8. Juni / 04 /ANDREAS / Gonsenheim” // “PARATYPUS 0 / Eumedonia eu- / medon mayencis / EI- TSCHBERGER & / STEINIGER, 1975” // “Coll. Bytins- ki Salz / Eing. Nr 20, 1960” // “ZMH 827551”; “Mainz / Mombach / 10.6.1922” // “Eumedon” // “PARATYPUS o / Eumedonia eu- | medon mayencis | ETYTSCHBERGER & / STEINIGER, 1975” // “Coll. Bytinski Salz / Eing. Nr 20, 1960” // “ZMH 827552”.
Type locality. Germany: “Deutschland, Rhein- land-Pfalz, Mainz-Mombach, Mainzer Sand”.
80. moenus Eitschberger & Steiniger, 1975
Original combination. “Eumedonia eumedon moenus n. subsp.” Eitschberger & Steiniger, 1975 Atalanta 6 (2): 99.
Current combination. Eumedonia eumedon moenus Eitschberger & Steiniger, 1975.
Current status. Valid subspecies.
Type material. Paratypes 2¢¢ (ZMH 827546-— 827547) (Fig. 80). “Tiertalsberg be1/ Retzbach am Main
Evolutionary Systematics 5 2021, 193-261
/ 4.6.1950” // “Katalog Nr. / L 3370” // “PARATYPUS o / Eumedonia eume- / don moenus | EITSCHBERG- ER & / STEINIGER, 1975” // “Bot.-Zool. Excursion / 1950/ Eing Nr. 14—1950” // “ZMH 827546”; “Karl- stadt a. Main / Karlsburg / 4.6.1950” // “Katalog Nr. / L 3369” // “PARATYPUS 0 / Eumedonia eume- / don moenus | EITSCHBERGER & / STEINIGER, 1975” // “Bot.-Zool. Excursion / 1950/ Eing Nr. 14—1950” // “ZMH 827547”.
Type locality. Germany: “Deutschland, Nordbayern, mittleres Maintal und Nord-Baden, mittleres Taubertal.”
81. persa Bytinski-Salz & Brandt, 1937
Original combination. “Chilades trochylus, Frr. ssp. persa, ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: Suppl. (2).
Current combination. Freyeria trochylus persa (By- tinski-Salz & Brandt, 1937).
Current status. Valid subspecies.
Type material. Syntypes 4¢¢ (ZMH 827587-— 827590) (Fig. 81). “IRAN / Keredj (1600 m.) / 3/6 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S + Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827587”; “IRAN / Keredj (1600 m.) / 9/8 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S + Brandt” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827588”; “IRAN / Keredj (1600 m.) / 12/8 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / By.S + Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827589”; “IRAN / Keredj (1600 m.) / 3/6 1936 / Brandt” // “Cotypus / Ch. trochylus / ssp. persa / ByS + Br.” // “Coll. Bytinski-Salz / Eing. Nr 20, 1960” // “ZMH 827590”.
Type locality. Iran: Tehran, “Keredj” [Karaj].
82. ab. pauper Bytinski-Salz & Brandt, 1937
Original combination. “Chilades trochylus, Frr. ssp. persa, ssp. nov. ab. pauper, ab. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: Suppl. (2).
Current combination. Freyeria trochylus persa ab. pauper (Bytinski-Salz & Brandt, 1937).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Holotype” 9 (ZMH 827591) (Fig. 82). “IRAN / Keredj (1600 m.) / 25/5 1936 / Brandt” // “ab. pauper / ByS+Br. / Holotype 9 / ex. coll. / Dr. H. Bytinski-Salz” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827591”.
Original locality. Iran: Tehran, “Keredj” [Karaj].
Remarks. Bytinski-Salz and Brandt (1937) proposed this name as an aberration of C. trochylus (Freyer, 1838). Therefore, as stated by article 45.6.2 ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
223
83. parvandereggi Verity, 1938
Original combination. “Glaucopsyche cyllarus, Rott, race parvandereggi, nom. nov.” Verity, 1938 Ent. Rec. 50: Suppl. (4).
Current combination. Glaucopsyche alexis parvan- dereggi Verity, 1938.
Current status. Valid subspecies.
Type material. Syntypes 246 (ZMH 833470- 833471) (Fig. 83). “parvandereggi Vrty” // “Macedonia / Olympus, 2900’ / V. 25, 1936 / Romei” // [blank label] // “ZMH 833470”; “parvandereggi Vrty” // “Macedonia / Olympus, 2900’ / V. 26, 1936 / Romei” // [blank label] // “ZMH 833471”.
Type locality. Greece: Salonika [Macedonia, Olympus].
Remarks. Verity (1938) proposed this name as a race of G. cyllarus (Rottemburg, 1775). As it was given in trinomy and the content of the description clearly indi- cates that a geographical race was meant, this name 1s fully available as subspecific (article 45.6 ICZN 1999). (Alberto Zilli pers. comm. ).
84. sibyllina Verity, 1916
Original combination. “Lycaena coridon” Poda, “razza sibyllina” Verity, 1916 Boll. Soc. Ent. Ital. 46: 133.
Current combination. Lysandra coridon sibyllina (Verity, 1916).
Current status. Junior subjective synonym of Lysan- dra coridon apennina (Zeller, 1847).
Original material. Labelled as “Type” 404 (ZMH 833493-833496) (Fig. 84). “sibyllina Vrty” // “Marche / Bolognola / 15 Ag 1937 / Querci” // [blank label] // “ZMH 833493”; “sibyllina Vrty” // “Marche / Bolognola/9 Ag 1937 / Querci” // [blank label] // “ZMH 833494”; “sibyllina Vrty” //““Marche / Bolognola / 9 Ag 1937 / Querci” // [blank label] /!“ZMH 833495”; “sibyllina Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 833496”.
Original locality. Italy: “Monti Sibillini (Bolognola, 1200 m)”
Remarks. Verity (1938) proposed this name as a “raz- za’ (=race) of L. coridon (Poda, 1761). As it was given in trinomy and the content of the description clearly indicates that a geographical race was meant, this name is avail- able as subspecific (article 45.6 ICZN 1999). However, it was later synonimized with L. coridon apennina (Zeller, 1847). The year on the specimen labels (1937) indicate specimens were collected 21 years after description (1916) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
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224 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Coll. Bytinghi Salz
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| ene. enn 2MH - 827589 ZMH 827591
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ZMH 833470
1cm
Plate 12. 73. Lycaena virgaureae emilianus f. quercii (Turati, 1923); 74. Agriades pheretiades andarabi (Forster, 1938); 75. Aricia anteros f. altera (Zullich, 1929); 76. Aricia anteros r. modicior Verity, 1938; 77. Plebejus idas uliginosa (Dannehl, 1921); 78. Plebejus argyrognomon balcanica (Zillich, 1929); 79. Eumedonia eumedon mayencis Eitschberger & Steiniger, 1975; 80. Eumedonia eum- edon moenus Eitschberger & Steiniger, 1975; 81. Freyeria trochylus persa (Bytinski-Salz & Brandt, 1937); 82. Freyeria trochylus persa ab. pauper (Bytinski-Salz & Brandt, 1937); 83. Glaucopsyche alexis parvandereggi Verity, 1938; 84. Lysandra coridon sibyl- lina (Verity, 1916). a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
85. f. lunulata Warnecke, 1942
Original combination. “Lycaena alcon F., 9, n. f. lunulata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.
Current combination. Phengaris alcon f. lunulata (Warnecke, 1942).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 12 (ZMH 827528) (Fig. 85). “Bahrenfeld / 28.7.08” // “Sig. S. War- necke / Eing. Nr. 5 1949” // “9 nf. unuata / Warn. Type / Iris 1942, 103” // “Katalog Nr. L 32 67” // “ZMH 827528”.
Original locality. Germany: Eidelstedt, Hamburg.
Remarks. Warnecke (1942) proposed this name as a form of “ZL. alcon F.” (sic, recte: [Denis & Schiffer- miller], 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambig- uously reveals that the name was proposed for an infra- subspecific entity, and is hence unavailable. The title of the paper (Warnecke, 1942) indicates that this form con- stitutes an aberration and is therefore infrasubspecific and unavailable. According to the description this form was only found in a single female.
86. f. selzeri Warnecke, 1924
Original combination. “Lycaena arionides Staudinger f. selzeri” Warnecke, 1924 Dt. Ent. Z. Iris 38: 149.
Current combination. Phengaris arionides f. selzeri (Warnecke, 1924).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827545) (Fig. 86). “Ussuri” // “F. selzeri Warn. / 3 Type” // “nf. selzeri Warn. / I. Iris 1924, 38. J. / p. 149, Taf. I bis. 5.” // “Type” // “Lyc. arionides - / n.f. selzeri Warn. / L.33” // “Katalog Nr. / L 3336” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827545”.
Original locality. Russia: “Ostsibirien, Amur-Gebiet, Ussuri”.
Remarks. Warnecke (1942) proposed this name as a form of L. arionides Staudinger, 1887. According to arti- cle 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was pro- posed for an infrasubspecific entity, and is hence unavail- able. According to the description this form is an individ- ual aberration or a rare second generation and is therefore infrasubspecific. Accordingly, on plate II this form is fig- ured as “Lycaena arionides nov. ab. selzeri Warn.”.
87. peninsulae Verity, 1923
Original combination. “Lycaena euphemus, Hub. (a) race peninsulae, mihi” Verity, 1923 Ent. Rec. 35 Suppl. (7).
Current combination. Phengaris euphemus penin- sulae (Verity, 1923).
225
Current status. Valid subspecies.
Original material. Labelled as “Type” 14 (ZMH 833468) (Fig. 87). “peninsulae Vrty” // “Marche / Bolog- nola / 10.VI.1937 / Querci / 1300 m” // [blank label] // “ZMH 833468”.
Original locality. Italy: “Sibillini Mts. in the Marche, below Bolognola, at 1000 m”.
Remarks. Verity (1923) proposed this name as a race of L. euphemus Hubner, 1799-1800. According to Kudrna (1983: 35), this race represents an available subspecific name. However, the year on the specimen labels (1937) indicate specimens were collected 14 years after descrip- tion (1923) from the same location (..e., Italy: Bolognola) the original type material was collected. As a result, the black label of the specimens treated as “type” by Veri- ty is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, this specimen is erroneously la- belled as type, should be annotated as “Not a Type” and are treated here as “Non-type” specimen.
88. ab. depuncta Hirschke, 1903
Original combination. “Lycaena Argiades ab. (et var.) depuncta” Hirschke, 1903 Verh. Ges. Wien. 53: 270.
Current combination. Cupido argiades ab. depunc- ta (Hirschke, 1903).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827531) (Fig. 88). “Hirschke.Wien / Herkulsbad / V.1900 Type” // “Verh. zool. / bot. ges. / 1903 Hirs.” // “TYPE” /1 “ZMH 827531”.
Original locality. Romania: “Herkulesbad”.
Remarks. Hirschke (1903) proposed this name as an aberration of L. argiades (Pallas, 1771). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
89. ausonia Verity, 1913
Original combination. “Lycaena damon ausonia nom. nov.” Verity, 1913 Boll. Soc. Ent. Ital. 45: 232, pl. 1, f. 44,47.
Current combination. Polyommatus (Agrodiaetus) damon ausonia (Verity, 1913).
Current status. Junior subjective synonym of Poly- ommatus (Agrodiaetus) damon damon ({Denis & Schif- fermuller], 1775).
Original material. Labelled as “Type” 63'¢ (ZMH 833479-833484) (Fig. 89). “ausonia Vrty.” // “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “ZMH 833479”; “ausonia Vrty.” // “Marche / Bolognola /
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226 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
15.VH.1937 / Querci” // [blank label] // “ZMH 833480”; “ausonia Vrty.” // “Marche / Bolognola / 12.VII.1937 / Querci” // [blank label] // “ZMH 833481”; “auso- nia Vrty.” // “Marche / Bolognola / 5 Ag 1937 / Quer- ci” // [blank label] // “ZMH 833482”; “ausonia Vrty.” // “Marche / Bolognola/3 Ag 1937 / Querci” // [blank label] //“ZMH 833483”; “ausonia Vrty.” // “Marche / Bolognola / 12. VII.1937 / Querc1” // [blank label] // “ZMH 833484”.
Original locality. Italy: “a grandi altitudini nei soliti monti del Piceno” [At high elevations at Mount del Piceno].
Remarks. The year on the specimen labels (1937) in- dicate specimens were collected 24 years after descrip- tion (1913) from the same location the original holotype material was collected. As a result, the black label of the Specimens treated as “types” is erroneous. The speci- men collected from the same geographical location from which the type specimen was collected is called topo- type. According to ICZN (1999), a topotype has no for- mal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
90. pfeifferi Brandt, 1938
Original combination. “Lycaena Pfeifferi sp. n.” Brandt, 1938 Ent. Rdsch. 55: 671, pl. 4, f. 1-3.
Current combination. Polyommatus (Agrodiaetus) Dfeifferi (Brandt, 1938).
Current status. Valid species.
Type material. Paratype 14 (ZMH 827579) (Fig. 90). “IRAN. FARS, StrafBe / Ardekan — Talochosroe / Comé, m, 1937 / coll. Brandt” //“L.37 / Paratypoid / Lyc. Dfeifferi Brand” // “Katalog Nr. / L. 3190” // “Sammlung R. Jahnig / Eing. Nr.29, 1949” // “ZMH 827579”.
Type locality. Iran: Fars “vom Barm I Firus (3750 m)” [from Barm-e Firuz].
91. alcestis Zerny, 1932
Original combination. “Lycaena (Hirsutina) ripperti alcestis Zerny nov. subsp.” Zerny, 1932 Dt. Ent. Z. Iris 46: 186.
Current combination. Polyommatus (Agrodiaetus) alcestis (Zerny, 1932).
Current status. Valid species.
Type material. Syntypes 2¢¢ (ZMH 827529- 827530) (Fig. 91). “Syria / Libanon s. / Bscharre / VIII- IX 31 / 1300 m / Einh.Slir.leg” // “Cotypus / ripperti / ssp. alcestis / Zerny” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // “ZMH 827529”; “Syria / Libanon s. / Bscharre / VIU-IX 31 / 1300 m/ Einh.Slrleg” // “Cotypus / ripperti / ssp. alcestis / Zerny” // “Coll. Bytinski-Salz / Eing Nr 20, 1960” // “ZMH 827530”.
Type locality. “Lebanon, Becharré, 1400 m; Zedern- wald oberhalb Becharré um 1900 m”.
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92. ardschira Brandt, 1938
Original combination. “Lycaena ardschira sp. n.” Brandt, 1938 Ent. Rdsch. 55: 672.
Current combination. Polyommatus (Agrodiaetus) ardschira (Brandt, 1938).
Current status. Valid species.
Type material. Paratype 1, (ZMH 827532) (Fig. 92). “L.36 / Paratypoid / Lyc. ardschira / Brand” // “Paratyp- oid” // “IRAN. FARS, StraBe / Ardekan — Talochosroe / Comé, m, 1937 /coll. Brandt” // “196” // “Sammlung / R. Jahnig / Eing. Nr. 29,1949” // “Katalog Nr. / L. 3365” // “ZMH 827532”.
Type locality. Iran: Fars, “Comée, 2600 m” [Komehr].
93. f. squalida Verity, 1920
Original combination. “Polyommatus meleager, Esp., race macra, mihi, 2 form squalida, mihi” Verity, 1920 Ent. Rec. 32:145.
Current combination. Polyommatus daphnis macra f. squalida Verity, 1920.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 24: (ZMH 833485—833486) (Fig. 93). “squalida Vrty” // “Marche / Bolognola / 15.VII.1937 / Querci” // [blank label] // “ZMH 833485”; “squalida Vrty” // “Marche / Bolognola /3 Ag 1937 / Querci” // [blank label] // “ZMH 833486”.
Original locality. Italy: “Mt. Sibillina, 1200-1300 m”.
Remarks. Verity (1920) proposed this name as a form of P. meleager macra Verity, 1920. According to Kudrna (1983: 39), this name is unavailable and denotes a race and is infrasubspecific and hence unavailable. Papilio meleager Esper, 1779 is a junior subjective synonym of Papilio daphnis {Denis & Schiffermiuller], 1775. The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geo- graphical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these two specimens are erroneous- ly labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
94. r. correpta Verity, 1920
Original combination. “Agriades hylas, Esp., race cor- repta, mihi” Verity, 1920 Ent. Rec. 32: 144.
Current combination. Polyommatus dorylas r. cor- repta (Verity, 1920).
Current status. Infrasubspecific and hence unavail- able name.
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Original material. Labelled as “Type” 40:40 (ZMH 833489-833492) (Fig. 94). “correpta Vrty” // “Marche / Bolognola / 15.VII.1937 / Querci / 1500 m” // [blank label] // “ZMH 833489”; “correpta Vrty” // “Marche / Bo- lognola / 14.VII.1937 / Querci / 1400 m” // [blank label] // “ZMH 833490”; “correpta Vrty” // “Marche / Bolog- nola / 29. VII.1937 / Querci / 1600 m” // [blank label] // “ZMH 833491”; “correpta Vrty” // “Marche / Bolognola / 12. VII.1937 / Querci / 1500 m” // [blank label] // “ZMH 833492”.
Original locality. Italy: “Bolognola, Central Italy”.
Remarks. Verity (1920) proposed this name as a race of A. hylas (Esper, 1779). According to Kudrna (1983: 26), this name is unavailable as it denotes a seasonal form which is infrasubspecific and hence unavailable. The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geo- graphical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneous- ly labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
95. olympena Verity, 1936
Original combination. “L. escheri, Hub. race olympena, nom. nov.” Verity, 1936 Ent. Rec. 48: 8.
Current combination. Polyommatus escheri olympe- na (Verity, 1936).
Current status. Valid subspecies.
Type material. Syntypes 246 (ZMH 833487-— 833488) (Fig. 95). “olympena Vrty” // “Macedonia / Olympus, 2500’ / July 13, 1935 / Romew” // [blank la- bel] // “ZMH 833487”; “olympena Vrty” // “Macedonia / Olympus, 2500’ / July 9, 1936 / Romei” // [blank label] /1 “ZMH 833488”.
Type locality. Greece: “Mt. Olympus, from 850 to 1500 m.”
Remarks. Verity (1936) proposed this name as a race of L. escheri Hubner, 1823. According to Kudrna (1983: 34), this race represents an available subspecific name.
96. ab. privata Schinfeld, 1924
Original combination. “Lycaena icarus” Rott.-3 “ab. privata m., nov. ab.” Schoenfeld, 1924 Int. Ent. Z. 18: 40.
Current combination. Polyommatus icarus ab. pri- vata (Schonfeld, 1924).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827573) (Fig. 96). “14.5. 1921 / KI. Chuchel / Bohmen /
227
von Schonfeld” // “Type / Lyc. icarus Rott. 3 / ab. privata Schonfeld /1I.E.Z. Guben. 18. Jahr. Nr 6/1924. a.d. nachl. v. Schonfeld. / D. Gelpke” // “KI — Chuchel / 14.5.21” / “ZMH 827573”.
Original locality. Czech Republic: “in der Umgebung von Prag (bei Kuchelbad)* [in the vicinity of Prague (near Kuchelbad)].
Remarks. Schoenfeld (1924) proposed this name as an aberration of L. icarus Rottemburg, 1775. Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
97. nuksani Forster, 1937
Original combination. “Lycaena candalus nuksani ssp. nov.” Forster, 1937 Mitt. Mtinch. Ent. Ges. 27: 62.
Current combination. Polyommatus (Sublysandra) nuksani (Forster, 1937).
Current status. Valid species.
Type material. Paratypes 2¢.¢ 29 9, (ZMH 827537-— 827540) (Fig. 97). “Nord-Ost-Hindukusch / Nuk- san-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. / W. Forster Munchen” // “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827537”; “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Al- penwiesenzone / 3500-4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus / nuksani Forst. / W. Forster Munchen” // “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827538”; “Nord-Ost-Hindukus- ch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500- 4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus | nuksani Forst. / W. Forster Miinchen” // “candalus / nuksani” // “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827539”; “Nord-Ost-Hindukus- ch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500- 4000 m Mitte Juli / leg. H.&E. Kotzsch” // “Para-Type / Lyc. candalus | nuksani Forst. / W. Forster Miinchen” // “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827540”.
Type locality. Afghanistan: “Nordost-Hindukusch, Nuksan-PaB Nordseite, 3500-4000 m.”.
98. ab. clorinda Verity, 1938
Original combination. “Tarucus balkanica, Freyer, with I gen. clorinda, nom. nov.” Verity, 1938 Ent. Rec. 50: (8).
Current combination. Tarucus balkanica ab. clorin- da Verity, 1938.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 24'3 (ZMH 835592-835593) (Fig. 98). “clorinda Vrty” // “Macedo- nia / Salonika, 1000’ / May 31, 1935 / Rome” // [blank label] // “ZMH 835592”; “clorinda Vrty” // “Macedonia /
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228 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Lye. arronides - Tr afr diac rane ce
[ZMH -827531|
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Plate 13. 85. Phengaris alcon f. lunulata (Warnecke, 1942); 86. Phengaris arionides f. selzeri (Warnecke, 1924); 87. Phengaris euphemus peninsulae (Verity, 1923); 88. Cupido argiades ab. depuncta (Hirschke, 1903); 89. Polyommatus (Agrodiaetus) da- mon ausonia (Verity, 1913); 90. Polyommatus (Agrodiaetus) pfeifferi (Brandt, 1938), 91. Polyvommatus (Agrodiaetus) alcestis (Zerny, 1932); 92. Polyommatus (Agrodiaetus) ardschira (Brandt, 1938); 93. Polyommatus daphnis macra f. squalida Verity, 1920; 94. Polyommatus dorylas tr. correpta (Verity, 1920). a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
Salonika, 1000’ / May 31, 1935 / Rome1” // [blank label] /1 “ZMH 835593”.
Original locality. Greece: “Salonika”.
Remarks. Verity (1938) proposed this name as an aberration of 7’ balkanica (Freyer, 1844). Therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
99. ahasveros Bytinski-Salz & Brandt, 1937
Original combination. “Lycaena (Glaucopsyche) pana- gaea, H.-Sch. ssp. ahasveros ssp. nov.” Bytinski-Salz & Brandt, 1937 Ent. Rec. 49: Suppl. (2).
Current combination. Turanana panagaea ah- asveros (Bytinski-Salz & Brandt, 1937).
Type material. Paratypes 1432 (ZMH 827556-— 827569) (Fig. 99). “IRAN: / Keredj (1700 m.) / 6.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “ssp. ahasveros / ByS + Brdt.” // “Coll. By- tinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827556”; “Tran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytins- ki-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827557”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratyp- oid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827558”; “IRAN: / Keredj (1200 m.) / 6.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827559”; “IRAN: / Keredj (1700 m.) / 6.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros / ByS+Brdt” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827560”; “Iran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / coll. Bytinski-Salz” //“Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827561”; “Iran: / Keredj 1700 m/ V.1936/ leg. F. Brandt / coll. Bytinski-Salz” // “Paraty- poid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827562”, “Iran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827563”; “Tran: / Keredj 1700 m/ V.1936/ leg. F. Brandt / coll. Bytinski-Salz” // “Paraty- poid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827564”; “Tran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827565”; “IRAN: / Keredj (1700 m.) / 10.5.1936 / Brandt” // “Cotypus / L. panagaea / ssp. ahasveros | ByS+Brdt” // “ssp. ahasveros / ByS + Brdt.” // “Coll. Bytinski-Salz / Eig. Nr. 20, 1960” // “ZMH 827566”; “Iran: / Keredj 1700 m/ V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827567”; “Iran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” /1 “ZMH 827568”; “X” // “Tran: / Keredj 1700 m / V.1936 / leg. F. Brandt / coll. Bytinski-Salz” //“Paratypoid” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827569”.
Type locality. Iran: Tehran, “Keredj 1700 m.” [Karaj].
Current status. Valid subspecies.
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Subfamily Theclinae Swainson, 1831 Tribe Eumaeini Doubleday, 1847
100. syra Pfeiffer, 1932
Original combination. “7hecla ilicis Esp. ssp. n. syra m.” Pfeiffer, 1932 in Osthelder and Pfeiffer 1932, Mitt. Munch. Ent. Ges. 22: 31.
Current combination. Satyrium ilicis syra (Pfeiffer, 1932).
Current status. Junior subjective synonym of nomi- notypical Satyrium ilicis (Esper, 1779).
Type material. Syntypes 34'\4 (ZMH 827570-827572) (Fig. 100). “Syria sept. / amantis / VI.1922” // “Cotypus / ilicis / ssp. syra/ Pfeiffer” // “ssp. syra/ Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 827570”; “Syria sept. / Taurus / Marasch / 26.V.28 / L. Osthelder leg. / 900 m” // “Cotypus / ilicis / ssp. syra / Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 827571”; “Syria sept. / Taurus / Marasch / 26.V.28/L. Osthelder leg. / 1200 m” // “Cotypus / ilicis / ssp. syra/ Pfeiffer” // “Coll. Bytinski-Salz Coll. / Eing. Nr. 20, 1960 Eing.” // “ZMH 827572”.
Type locality. Turkey: “Syria sept., Taurus, Marasch” [Marash].
101. brevicaudis Vorbrodt, 1911
Original combination. “Thecla linceus Esp. (= spini Schiff.) brevicaudis Pingeler, 1.1.” Vorbrodt, 1911 Die Schmetterlinge der Schweiz. I.: 106.
Current combination. Satyrium spini brevicaudis (Vorbrodt, 1911).
Current status. Junior subjective synonym of nomi- notypical Satyrium spini ({[Denis & Schiffermiuller], 1775).
Type material. Syntype | 2 (ZMH 827574) (Fig. 101). “Zermatt / Rpe an Rhamnus alp.[ina] / myrmecophil mit / Formica fusca 6.1908 / 8 cl. 28.7.1908 Ping.” // “Co- type / v. brevicaudis Ping.” // “Spini var. brevicaudis Pung” // “L.29” // “Katalog Nr. / L 2048” // Sig. G. “War- necke / Eing. Nr. 5, 1949” // “ZMH 827574”.
Type locality. Switzerland: “bei Zermatt 1700 m”.
Remarks. Vorbrodt (1911) proposed this name as a form (in the original description and “var.” on the spec- imen labels) of “7! Linceus” (sic, recte: [ynceus) (Esper, 1779). According to article 45.6.4 (ICZN 1999), it is sub- specific if first published before 1961 and author express- ly used the term “form” or “var”.
Unpublished, unavailable names
102. v. romana Hartig
Original combination. Lycaena cyllarus v. romana Hartig.
Current status. “in litteris’” name and hence not available.
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230 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Original material. Labelled as “Cotype” 3¢.¢ (ZMH 833472-833474) (Fig. 102). “romana Hartig” // Lazio / Formia / 23 4 [19]38 / Querci” // [blank label] // “ZMH 833472”; “romana Hartig” // Lazio / Formia / 8 4 [19]38 / Querci” // [blank label] // “ZMH 833473”; “romana Har- tig” // Lazio / Formia / 3 5 [19]38 / Querci” // [blank la- bel] // “ZMH 833474”.
Original locality. Italy: Lazio.
Remarks. This taxon cannot be found in the literature. However, it may refer to Glaucopsyche alexis latina Har- tig, 1940 (Mem. Soc. Ent. Ital. 18: 187; Type locality: Formia, Latina, Italy). The label data matches the type locality of /atina. Papilio cyllarus Rottemburg, 1775 is a junior subjective synonym of Papilio alexis Poda, 1761.
103. ab. albipunctata Hartig
Original combination. Lysandra coridon ab. albipunc- tata Hartig.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 446 (ZMH 833475833478) (Fig. 103). “Typen” // “19/7 1917 / Gottingen / Hohefeld” // “ZMH 833475”; “13/7 1917 / Gottingen / Hohefeld” // “ZMH833476”; “18/8 1917 / Gottingen / Hohefeld” // “ZMH833477”; “18/7 1917 / Gottingen / Hohefeld” // “ZMH 833478”.
Original locality. Germany: Gottingen.
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
Family Nymphalidae Rafinesque, 1815 Subfamily Danainae Boisduval, 1833
104. icensis Fuchs, 1954
Original combination. “Danais gilippus var. icensis” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.
Current combination. Danaus gilippus_ icensis (Fuchs, 1954).
Current status. Junior subjective synonym of Danaus eresimus erginus (Godman & Salvin, 1897).
Type material. Lectotype 14 (ZMH 827645), para- lectotypes 2346 (ZMH 827643-827644) (Fig. 104). “Huayuri / 5/1940 / Type” / “Coll. H. Rodinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “ZMH 827643”; “Ica / 7/1940 / Type” / “Coll. H. Rodinger / Eing. Nr. 30, 1952 / Katalog Nr. L” //“Paralectotype 9 / Danais gilippus / vat. icensis / von Fuchs / G. Lamas ‘98” // “Danaus eresimus / erginus Godman + / Salvin 2 / G. Lamas. Det. 1978” // “ZMH 827644”; “114 / eresimus. ab / Ica (Pert)” // “Ica / 7.1928 / Type” // “Coll. H. Rodinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “Lectotype 3 / Danais gilippus / var. icensis / von Fuchs / G. Lamas *78” // “Danaus eresimus / erginus Godman + / Salvin ¢ / G. Lamas. Det. 1978” // “ZMH 827645”.
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Type locality. Peru: “Im Tal von Ica und des Rio Grande”.
Remarks. Fuchs (1954) proposed this name as a “var.” of Danais gilippus Cramer, 1775. According to article 45.6.4 (ICZN 1999), it is subspecific if first published be- fore 1961 and author expressly used the term “var.”.
105. roedingeri Fuchs, 1954
Original combination. “Danais gilippus var. Rodingeri” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.
Current combination. Danaus gilippus roedingeri (Fuchs, 1954).
Current status. Junior subjective synonym of D. g. candidus Clark, 1941.
Type material. Holotype 1 (ZMH 827652), paratypes 63:2 (ZMH 827649-827651, ZMH 827653-827655) (Fig. 105). “Danais / gilippus / var. rédingeri & / v. Fuchs” // “Type” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “Std — Peru / Ayacucho — Yucaes / 2500 m 26. 4. 1936.” // “Holotype 3 / Danais gilippus / var. Rédingeri / von Fuchs” // “Danais gilippus 3 / candidus Clark / G. Lamas. det. 1978” // “Hbg. — Stid — Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” // “ZMH 827652”; “Ayacucho / 2—3000 m / 8/1920 / Type” // “D. rédingeri 2” // “Coll. H. Rédinger / Eing. Nr. 30, 1952” // “ZMH 827649”; “Type” // “Danais / gilippus // var. rédingeri & / v. Fuchs / Rio Jucaes b/ / Aya- cucho / 1800-2300” //“Summlung / W. v. Fuchs / Eng. Nr. 4, 1956” //“ZMH 827650”; “Ayacucho / 2—3000 m/ 8/1920 / Type” // “Coll. H. Rédinger / Eing. Nr. 30, 1952 / Katalog Nr. L” // “ZMH 827651”; “Ayacucho 28.4” // “Stid — Peru / Oberh. Ayacucho / 29.4.1936.” // “Ayacucho / 2—3000 m / 8/1920 / Type” // “Summlung / W. v. Fuchs / Eing. Nr. 4, 1956” //“Hbg. — Sud— Peru /Sammelreise 1936. / Eing. Nr. 1, 1937.” // “ZMH 827653”; “Danais / gilippus / var. r6din- geri 2 / v. Fuchs” // “Type” // “Sammlung / W. v. Fuchs / Eig. Nr. 4, 1956” // “Std — Peru / Ayacucho — Yucaes / 2500 m 26. 4. 1936.” // “Hbg. — Sid — Peru / Sammelreise 1936. / Eing. Nr. 1, 1937.” //“Danais gilippus 9 / candidus Clark /G. Lamas. det. 1978” // “Paratype / Danaus gilippus 9 / var. Rédingeri / von Fuchs” // “ZMH 827654”, “Type” // Danais | gilippus / var. rédingeri ° / v. Fuchs / Rio Jucaes / 6/ Ayacucho / 1800—2300” // “ZMH 827655”.
Type locality. Peru: “... naheren Umgebung von Ay- acucho, auf 2000-2600 m” [vicinity of Ayacucho, 2000- 2600 m].
Remarks. Fuchs (1954) proposed this name as a “var.” of Danais gilippus (Cramer, 1775). According to article 45.6.4 (ICZN 1999), it is subspecific if first published be- fore 1961 and author expressly used the term “var.”.
Subfamily Heliconiinae Swainson, 1822
106. altonevadensis Reisser, 1927
Original combination. “Argynnis niobe var. nov. altone- vadensis” Reisser, 1927 Int. Ent. Z. Guben 20: 373.
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W- chesckel 4S. 4
Macedonia ‘Sal sea =
935 tas homme
4
Goll. Bytinski-Salz. Eing, Nr. 20 - 62.
oT ay
Colt, Bytinski-Szl2Coli, Eing. Nr. 26, 1960Eing.
ZMH 827570
— 0 7
:
[ZMH 833473
[zm 833478
Plate 14. 95. Polyommatus escheri olympena (Verity, 1936); 96. Polyommatus icarus ab. privata (Schonfeld, 1924); 97. Polyommatus (Sublysandra) nuksani (Forster, 1937); 98. Tarucus balkanica ab. clorinda Verity, 1938; 99. Turanana panagaea ahasveros (Bytins- ki-Salz & Brandt, 1937); 100. Satyrium ilicis syra (Pfeiffer, 1932), 101. Satyrium spini brevicaudis (Vorbrodt, 1911); 102. Lycaena cyllarus v. romana Hartig; 103. Lysandra coridon ab. albipunctata Hartig. a. Dorsal view, b. ventral view.
1 cm
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232 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Current combination. Fabriciana niobe altone- vadensis (Reisser, 1927).
Current status. Junior subjective synonym of Fabri- ciana niobe niobe (Linnaeus, 1758).
Type material. Syntypes 1 ¢ 192 (ZMH 827656- 827657) (Fig. 106). “Type # 7 3 / Argynnis niobe var. / altonevadensis Reisser / Int. Ent. Z. guben / Summer 1927” // “HISPANIA VII 26 / SIERRA NEVADA / O BUBACEK” // [blank label] // “ZMH 827656”; “Type # 4 9 / Argynnis niobe var. / altonevadensis Reisser / Int. Ent. Z. guben / Summer 1927” // “HISPANIA VII 26 / SIERRA NEVADA / O BUBACEK” // [blank label] // “ZMH 827657”.
Type locality. Spain: “... auf den Hohen der Sierra Nevada in Stidspanien ...“. [higher areas of Sierra Neva- da in southern Spain].
Remarks. Reisser (1927) proposed this name as a “var.” of Argynnia niobe (Linnaeus, 1758). According to article 45.6.4 (ICZN 1999), it is subspecific if first published be- fore 1961 and author expressly used the term “var.”.
(<4
107. cyrnea Schawerda, 1926
Original combination. “Argynnis pandora Schiff. var. nov. cyrnea” Schawerda, 1926 Z. Ost. Ent. Ver. 11: 109.
Current combination. Argynnis pandora cyrnea Schawerda, 1926.
Current status. Junior subjective synonym of Argy- nnis pandora pandora ({Denis & Schiffermiller], 1775).
Type material. Syntypes 2¢¢ (ZMH 833447- 833448) (Fig. 107). “pandora / var. cyrnea Schawerda / Cotype [illegible]” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 833447”; “pandora / var. cyrnea Schawerda / Cotype [illegible]” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 833448”.
Type locality. France: “Korsika” [Corsica].
Remarks. Schawerda (1926) proposed this name as a “var.” of Argynnis pandora ([Denis & Schiffermiuller], 1775). According to article 45.6.4 ICZN 1999), it is sub- specific if first published before 1961 and author express- ly used the term “var.”.
108. argentifasciata Kotzsch, 1937
Original combination. “Argynnis pandora Schiff. ar- gentifasciata ssp. n.” Kotzsch, 1937 Ent. Rdsch. 55: 9.
Current combination. Argynnis pandora argentifas- ciata Kotzsch, 1937.
Current status. Valid subspecies.
Type material. Syntypes 24.4 (ZMH 827665-827666) (Fig. 108). “West-Hindukusch / Andarab / Gerollzone / 4000-4500 m Mitte August / leg. H.&E. Kotzsch” // “Co- typus / A. pandora / ssp. argenti- / fasciata / Kotzsch” // “ssp. argentifasciata / Kotzsch” // “ZMH827665”; “West-Hindukusch / Andarab / Banu-Ebene / 2000-
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2500 m August /leg. H.&E. Kotzsch” //“Cotypus / A. pan- dora/ ssp. argenti- / fasciata / Kotzsch” //“ZMH827666”.
Type locality. Afghanistan: “Andarab im westlichen Hindukusch”.
Subfamily Nymphalinae Rafinesque, 1815 Tribe Melitaeini Newman, 1870
109. f. urbanoides Warnecke, 1942
Original combination. “Melitaea maturna L., 3, n. f. urbanoides.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.
Current combination. Euphydryas maturna f. ur- banoides (Warnecke, 1942).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 24:3 (ZMH 827658-827659) (Fig. 109). “Holstein / Innien, el. / 1913 Selzer” // “3 nf. urbanoides / Warn. Type / Iris 1942, 103” // “L.25 / Melitaea maturna - / urbanoides / Warn.” // “ZMH 827658”; “Holstein / Innien / [illegible]” /1 “VON / G. WARNECKE / W. LINZ. HAMB.” // “Sig. W. Linz / Eing. 12 — 74” // “ZMH 827658”.
Original locality. Germany: “Innien bei Neumutnster (Holstein).”; Schleswig-Holstein.
Remarks. Warnecke (1942) proposed this name as a “form” of Melitaea maturna (Linnaeus, 1761). Accord- ing to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. The title already indicates that this form constitutes an individual aberration and thus an un- available name.
110. tenuicula Verity, 1919
Original combination. “Melitaea athalia race tenuicula mihi” Verity, 1919 Ent. Rec. 31: 193-194.
Current combination. Melitaea athalia tenuicula Verity, 1919.
Current status. Valid subspecies.
Original material. Labelled as “Type” 64° (ZMH 833455—-833460) (Fig. 110). “tenuicula Vrty” // helvet- ica Ruhl” // “Marche / Bolognola / 28 VI 1937 / Quer- ci.” // [blank label] // “ZMH 833455”; “tenuicula Vrty” // helvetica Ruhl” // “Marche / Bolognola / 15 VI 1937 / Querci.” // [blank label] // “ZMH 833456”; “tenuicula Vrty” // “Marche / Bolognola / 17 VI 1937 / Querci.” // [blank label] // “ZMH 833457”; “tenuicula Vrty” // hel- vetica Ruhl” // “Marche / Bolognola / 18 VI 1937 / Quer- ci.” // [blank label] // “ZMH 833458”; “tenuicula Vrty” // “Marche / Bolognola / 28 VI 1937 / Querci.” // [blank la- bel] // “ZMH 833459”; “tenuicula Vrty” // “Marche / Bo- lognola / 27 VI 1937 / Querci.” // [blank label] // “ZMH 833460”.
Original locality. Italy: “Bolognola in the Sibillini Mountains, 1200 m”.
Evolutionary Systematics 5 2021, 193-261 233
: Coll. H,.Rédinger : Eing. Nr. 30, 1952 ‘Katalog Nr. L
Se Danaus eresim us 3 Jabvin G.Lamas. det. 1976
Z7MH_827645|
Yerrscr¢ i
haa: 9¢Lf ee ohn vedenaered Ks ar
hace ‘ae Danaiy 9 lippus yer. pack wey
G. Lamas. det. 1978
F ~~ Sammlung a W. v. Fuchs :
Evisa_ Corse Jul 1921 Col.O,.Bubacek
=
HISPANIA VII 26 SIERRA. NEVADA Q BUBACEK
a
ZMH_827656
107b-
West-Hindukusch @ Andarab Geridlzone 4000-4500m Mitte August , leg. H.&E.Kotzsch
[ZMH 827665
IZMH 827658|
11cm
Plate 15. 104. Danaus gilippus icensis (Fuchs, 1954); 105. Danaus gilippus roedingeri (Fuchs, 1954); 106. Fabriciana niobe alto- nevadensis (Reisser, 1927); 107. Argynnis pandora cyrnea Schawerda, 1926; 108. Argynnis pandora argentifasciata Kotzsch, 1937; 109. Euphydryas maturna f. urbanoides (Warnecke, 1942). a. Dorsal view, b. ventral view.
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234 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Remarks. Verity (1919) proposed this name as a “race” of Melitaea athalia (Rottemburg, 1775). According to Kudrna (1983: 83-84), this name is available as subspecif- ic. The year on the specimen labels (1937) indicate spec- imens were collected 18 years after description (1919) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and 1s not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens
111. varissima Verity, 1914
Original combination. “MVelitaea [parthenie Bkh] varia Meyer-Dur varissima nom. nov.” Verity, 1914 (1913) Boll. Soc. Ent. Ital. 45: 210.
Current combination. Melitaea varia varissima Ver- ity, 1914.
Current status. Valid subspecies.
Original material. Labelled as “Type” 63'¢ (ZMH 833461—833466) (Fig. 111). “varissima Vrty” // “varia Meyer-Dir” // “Marche / Bolognola / 25.VI.1937 / Quer- ci” // [blank label] // “ZMH 833461”; “varissima Vrty” // “Marche / Bolognola / 20. V1.1937 / Querci” // [blank label] /!“ZMH 833462”; “varissima Vrty” // “Marche / Bologno- la/14.V1.1937 / Querci” // [blank label] // “ZMH 833462”; “yarissima Vrty” // “Marche / Bolognola / 14.VI.1937 / Querci” // [blank label] // “ZMH 833463”; “varissi- ma Vrty” // “Marche / Bolognola / 5.VI.1937 / Querci” // [blank label] // “ZMH 833464”; “varissima Vrty” // “Marche / Bolognola / 22. V1.1937 / Querci” // [blank label] /!“ZMH 833465”; “varissima Vrty” // “Marche / Bologno- la /25.V1.1937 / Querci” // [blank label] // “ZMH 833466”.
Original locality. Italy: Bologna “Nei Faggeti di Bologna, nella localita Costarotara e a Piano Astore dei Monti Sibillini, da 1200 a 1600 m. d’altitudine” [Bolo- gna, vicinity of Costarotara and Piano Astore of the Sibil- lini Mountains, 1200-1600 m.].
Remarks. Verity (1919) proposed this name as a “race” of Melitaea varia Meyer-Dir, 1851. According to Kudrna (1983: 84), this name is available as subspecific. The year on the specimen labels (1937) indicate specimens were collected 23 years after description (1914) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collected from the same geo- graphical location from which the type specimen was col- lected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non-type” specimens.
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112. riloensis Ziillich, 1936
Original combination. “Melitaea cinxia L. var. riloensis Ziillich.” Zullich, 1936 Z. Ost. Ent. Ver. 21: 27.
Current combination. Melitaea cinxia riloensis Ziil- lich, 1936.
Current status. Junior subjective synonym of Meli- taea cinxia Clarissa Staudinger.
Type material. Syntypes 2¢¢ (ZMH 827646- 827647) (Fig. 112). “COTYPE / Mel. cinxia L. / 3 v. ri- loensis / Z. d. O. Ent. V. 1936 / DR. ZULLICH, WIEN” // “BULGARIEN JUNI 1923 / RYLA GEB. 1200 bis 1800 m/O. BUBACEK WIEN” // [blank label] // “ZMH 827646”; “COTYPE / Mel. cinxia L. / 3 v. riloensis / Z. d. O. Ent. V. 1936 / DR. ZULLICH, WIEN” // “BUL- GARIEN JUNI 1923 / RYLA GEB. 1200 bis 1800 m / O. BUBACEK WIEN” // [blank label] // “ZMH 827647”.
Type locality. Bulgaria: “auf dem Steilhang ostlich des Riloklosters in einer Hohe von etwa 1200 m.” [at a steep slope east of the Rilo monastry at an altitude of about 1200 m]. 1901.
Remarks. Zillich (1936) proposed this name as a “var.” of M. cinxia (Linnaeus, 1758). According to arti- cle 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.
113. ab. mediofasciata Bubacek, 1926
Original combination. “Melitaea dejone H.G. ab. medi- ofasciata Bub. ab. nova.” Bubacek, 1926 Verh. Zool.-Bot. Ges. Wien 74—75: (9).
Current combination. Melitaea deione ab. medio- fasciata Bubacek, 1926.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 833454) (Fig. 113). “Type / Mel. rondoui / ab. mediofas- ciata / Otto Bubacek” // “France, Gedre / Hautes Pyrenées / Juli 1923 / Col. Otto Bubacek” // [blank label] // “ZMH 833454”.
Original locality. France: Pyrenées.
Remarks. Bubacek (1926) proposed this name as an aberration of M. dejone Geyer, 1832. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
114. ab. johni Stauder, 1922
Original combination. “M/. dictynna ab. johni” Stauder, 1922 Ent. Anz. 2: 44.
Current combination. Melitaea diamina ab. johni Stauder, 1922.
Current status. Infrasubspecific and hence unavail- able name.
Evolutionary Systematics 5 2021, 193-261
Original material. Labelled as “Type” 14 (ZMH 827648) (Fig. 114). “Lainau 800 m / [illegible] / 1.VII.17 Schawerda” // “Type” // “dictynna f. / johni Stauder” // “Type” // “ZMH 827648”.
Original locality. Austria: “Salzkammergut (Lainau- tale, Hallstatt)”.
Remarks. Stauder (1922) proposed this name as an aberration of M. dictynna Esper, 1778. As stated by arti- cle 45.6.2 (ICZN 1999) it is deemed to be an infrasubspe- cific name (the author used “aberration”, “ab.”) and is hence unavailable.
115. balcanica Ziillich, 1936
Original combination. “MVelitaea trivia Schiff. var. bal- canica Zullich.” Zullich, 1936 Z. Ost. Ent. Ver. 21: 27.
Current combination. Melitaea trivia balcanica Ziillich, 1936.
Current status. Junior subjective synonym of Meli- taea trivia lathon Fruhstorfer, 1917.
Type material. Syntypes 246 (ZMH 827663- 827664) (Fig. 115). “BULGARIEN JUNI 1928 / RYLA GEB. 1200 bis 1800 m/O. BUBACEKR” // [blank label] // “COTYPE / Mel. trivia/ Gv. balcanica / Z. d. O. Ent. V. 1936 /DR. ZULLICH, WIEN” // “ZMH 827663”; “BUL- GARIEN JUNI 1928 /RYLA GEB. 1200 bis 1800 m/O. BUBACEK” // [blank label] // “COTYPE / Mel. trivia / 3 v. balcanica / Z. d. O. Ent. V. 1936 / DR. ZULLICH, WIEN” // “ZMH 827664”.
Type locality. Bulgaria: “in den gebirgigen Teilen der zentralen und 6stlichen Balkanhalbinsel (Krain, Bosnien, Herzegowina etc.)”. [in the mountainous parts of the cen- tral and eastern Balkan peninsula].
Remarks. Zillich (1936) proposed this name as a va- riety of M. trivia ([Denis & Schiffermitller], 1775). Ac- cording to article 45.6.4 (ICZN 1999), it is subspecific if first published before 1961 and author expressly used the term “var.”.
116. catapelioides Stauder, 1918
Original combination. “Velitaea trivia catapelioides nov. subspec. m.” Stauder, 1918 Zeit. Wiss. Ins. Biol. 14: 57. Current combination. Melitaea trivia catapelioides Stauder, 1918. Current status. Junior subjective synonym of nomino- typical Melitaea trivia ({[Denis & Schiffermtller], 1775). Original material. Labelled as “Type” 14 (ZMH 827662) (Fig. 116). “Type” // “Luen / Calabria mer. / Aspromonte 950 m. / 6.VII 1920 / H. Stauder legit.” // “trivia catape - / lioides Stauder” // “Type” // “Sig. G. Warnecke / Eing. Nr. 5, 1945” // “ZMH 827662”. Original locality. Italy: “Unteritalien (Kalabrien)”. Remarks. The year on the specimen label (1920) indicates specimen was collected two years after
230
description (1918) from the same location (..e., Italy, Kalabrien) the original holotype material was collected. As a result, the label of the specimen treated as “Type” is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, this specimen is erroneously labelled as type, should be annotated as “Not a Type” and is treated here as “Non-type” specimen.
117. ignasiti de Sagarra, 1926
Original combination. “Melitaea trivia Schiff., rassa ignasiti, nova rassa” de Sagarra, 1926 Butl. Inst. Catal. Hist. Nat. (2)6: 131.
Current combination. Melitaea trivia ignasiti de Sagarra, 1926.
Current status. Valid subspecies.
Type material. Paralectotypes 1419 (ZMH 827660-— 827661) (Fig. 117). “CATALONIA / Seva 700 m. / 3.8.25 Romei” // “M. trivia — ignasiti Sag. / co-type 4 Querci” /! “ZMH 827660”; “CATALONIA / Seva 700 m. / 3.8.25 Romei” // “Sammlung / R. Jahnig / Eing. Nr.29,1949” // “M. trivia — ignasiti Sag. / co-type 2 Querci” // “L.28” // “ZMH 827661”.
Type locality. Spain: Catalonia, “Martorell”.
Remarks. de Sagarra (1926) proposed this name as a “rassa = race” of M. trivia ([Denis & Schiffermullar], 1775). The term “rassa = race” which was employed by de Sagarra in 1926 was often used in the current sense of subspecies. As a result, the first line of article 45.6 (ICZN 1999) — the rank denoted by a species-group name following a binomen is subspecific — is applied here, it is thus available as subspecific. Higgins (1941) synonymised it with M. trivia pseudodidyma Rebel, 1905 but he reconsidered it as a valid subspecies in his subsequent books. Garcia-Barros et al. (2013) made a comment in Fauna Ibérica that “the orange color of the Iberian specimens is lighter than those from other European areas; those have been attributed to subspecies “ignasiti de Sagarra, 1926”, which, however, does not seem significantly different. Furthermore, it does not seem significantly differentiated from a genetic point of view (Wahlberg pers. comm.).” The taxon ignasiti was reinstated as a valid species by van Oorschot and Coutsis (2014), and this was accepted by Macia et al. (2015) who designated lectotype and paralectotypes. Therefore, specimens at ZMH are by default paralectotypes. However, most other recent authors (e.g., Kudrna 2019) treat this taxon as a synonym to M. trivia, because there is no evidence for species status. Even Oorschot and Coutsis (2014) mention intermediates indicating hybridisation. It should be noted that the species name is not mentioned in the latest European checklist (Wiemers et al. 2018).
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236 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
BS . ) ee I ub aia 4 a rr Leinwicate \Ivta AMA Mitek Nin iy ea A. <7 GP he Belated gaan ash
Marche Bolognola LE wyP1987
uercl,
Wdvahue Ral ee
wlth hetediact. ..... Ab, medtolasciaty. 7] a acche
COTYPE Me aural,
France, Gédre Hautes Pyrenées Juli 1923 Col. Otto Bubacek
Tu, uloonsts Z.4.b6-EntV. 1936
DR. ZULLICH, WIEN
BULGARIEN JUN] 1523 RYLA GES. 1200 bes 1800 m Q. BUBACEK WIEN
[ZMH 833454!
BULGARIEN ..JUNI 1928 RYLA GEB. 1200 bis 1800 m 0. BUBACEK WIEN
COTYPE Md. tuvia Ov. baleanica 2ST Ent Ve 1936
DR. ZOLIICH, WIEN
ZMH 827663
CATALONIA Seva 700 m. 4.8.25 Romei
Calabria mer.
Asprofaonte m, OV77 Leas H. Stauder legit.
[ZMH _827660|
Sig. G. Warnacke Elng. Nr.5 ,1948
ZMH _ 827662
Plate 16. 110. Melitaea athalia tenuicula Verity, 1919; 111. Melitaea varia varissima Verity, 1914; 112. Melitaea cinxia riloensis Ziullich, 1936; 113. Melitaea deione ab. mediofasciata Bubacek, 1926; 114. Melitaea diamina ab. johni Stauder, 1922; 115. Melitaea trivia balcanica Zillich, 1936; 116. Melitaea trivia catapelioides Stauder, 1918; 117. Melitaea trivia ignasiti de Sagarra, 1926. a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261 Tribe Nymphalini Rafinesque, 1815
118. guhni Tschauner, 1926
Original combination. “Vanessa urticae var. guhni” Tschauner, 1926 Int. Ent. Z. 20: 229, 273.
Current combination. Aglais urticae (Tschauner, 1926).
Current status. Junior subjective synonym of Ag/ais urticae urticae (Linnaeus, 1758).
Type material. Paratype 1 (ZMH 827667) (Fig. 118). “Paratypoid” // “Vanessa urticae / nov. var. guhni / Para- type e. coll. / Tschauner, Berlin 1926” // “Coll. Bytins- ki-Salz / Eing. Nr. 20, 1960” // “ZMH 827667”.
Type locality. Germany: Berlin.
Remarks. This case seems ambiguous, because Tschauner (1926) writes that the new name constitutes a local variation, if not a new race, although he also notes that he is not sure whether the variation is really heritable. On page 273 he names it again as “Vanessa urticae var. Guhni nov. spec.“ [sic!]. See also the critical remark by Kitt (1927).
guhni
119. f. clarior Warnecke, 1959
Original combination. “Araschnia levana (Linnaeus) gen. vern. n. f. clarior’ Warnecke, 1959 Mitt. Faun. Arb. Gem. Schlesw. Holst. 1 (10.): 53.
Current combination. Araschnia levana f. clarior Warnecke, 1959.
Current status. Infrasubspecific and thus unavail- able name.
Original material. Labelled as “Type” 26’ (ZMH 825009-825010) (Fig. 119). “Schleswig / Elsdorfer Sch. / 20.3.97” // “n.f. clarior W. / Coll. Warnecke / Eing. Nr. 5-1949” // “ZMH 825009”; “Schleswig / Elsdorfer Sch. /c. 1. 20.3.97” // “nf. clarior W. / Coll. Warnecke / Eing. Nr. 5-1949” // “ZMH 825010”.
Original locality. Germany: Schleswig-Holstein.
Remarks. Warnecke (1959) proposed this name as an aberration of A. /evana (Linnaeus, 1758). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and is hence unavailable. From the text of the description it is unam- biguously clear that the author is describing an aberration and not a subspecies.
120. ab. marginelineata Horch, 1932
Original combination. “Araschnia levana L. gen. aest. prorsaab. marginelineata” Horch, 1932 Int. Ent. Z. 26:338. Current combination. Araschnia levana prorsa ab. marginelineata Horch, 1932. Current status. Infrasubspecific and hence unavail- able name.
237
Original material. Labelled as “Kotype” 14 (ZMH 825011) (Fig. 120). “f. marginelineata / Horch / Gub. I. E Z; 26., 1932/3, / P. 337.” // “Umgeg. v. Hamburg / Rad- bruch / e. |. 10.7.32 / R. Horch Hamburg” // “Kotype” // “ZMH 825011”.
Original locality. “Luneburger Heide”, Hamburg [Germany].
Remarks. Horch (1932) proposed this name as an abberation of A. /evana (Linnaeus, 1758). This species is known for having two forms, /evana and prorsa that represent the spring and summer broods respectively. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
Subfamily Satyrinae Boisduval, 1833 Tribe Satyrini Boisduval, 1833
121. alphea Warnecke, 1919
Original combination. “Satyrus arethusa Schiff. nov. subsp. alphea Warn.” Warnecke, 1919 Int. Ent. Z. 12: 182.
Current combination. Arethusana arethusa alphea (Warnecke, 1919).
Current status. Junior subjective synonym of Arethu- sana arethusa arethusa ({Denis & Schiffermiuller], 1775).
Type material. Syntype 13 (ZMH 827676) (Fig. 121). “Type” // nov. v. Alphea” / Warn.” // “Type” // “Griechen- Id.” // “ZMH 827676”.
Type locality. Greece: “Griechenland, Velucci und Parnass”’.
122. asperomontana Stauder, 1921
Original combination. “Satyrus circe L. subsp. n. asper- omontana” Stauder, 1921 Dt. Ent. Z. Iris 35: 28.
Current combination. Brintesia circe asperomon- tana (Stauder, 1921).
Current status. Junior subjective synonym of Brinte- sia circe circe (Fabricius, 1775).
Type material. Syntype 13 (ZMH 827680) (Fig. 122). “1/7 / Calabria mer. / Asperomonte 1200 m. / Polsi 1920 / H. Stauder legit.” // “Type” // “circe / asperomontana / Type” // “ZMH 827680”.
Type locality. Italy: “Polsibeken, Aspromonte 1350- 1500 m.”
123. f. rufobrunnea Warnecke, 1942
Original combination. “Coenonympha arcania L. n. f. rufobrunnea.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102. Current combination. Coenonympha arcania f. ru- fobrunnea Warnecke, 1942. Current status. Infrasubspecific and hence unavail- able name.
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238 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
SAL, a1’; a erin VY uv e ‘ ty!
mia ~ iy ae
ae See be Coli. Pytinski-Saiz ; ZMH 827667
' Coli. Warnecke fing. Nr §-1948
f | Vs aa Of uta le
Kg ae 46. 14343 ft 992 ¢
Umgeg.v-Hamburg bruce
Rad h C4732,
R.Horch Hamburg:
ZMH 827680
1cm
Plate 17. 118. Aglais urticae guhni (Tschauner, 1926); 119. Araschnia levana f. clarior Warnecke, 1959; 120. Araschnia levana prorsa ab. marginelineata Horch, 1932; 121. Arethusana arethusa alphea (Warnecke, 1919); 122. Brintesia circe asperomontana (Stauder, 1921). a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
Original material. Labelled as “Type” 1¢ (ZMH 827677) (Fig. 123). “Itzehoe / (Holstein)” // “n.f. rufobrun- nea Warn. / Type / Iris 1942, p. 102” // “Katalog Nr. 519 / HL” // “Coenon. arcania - / f. rufobrunnea / Warn. / L.38” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827677”.
Original locality. Germany: Schleswig-Holstein, Itzehoe.
Remarks. Warnecke (1942) proposed this name as a form of C. arcania (Linnaeus, 1761). According to article 45.6.1 ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was pro- posed for an infrasubspecific entity, and is hence unavail- able. From the text it is clear that the author describes an individual form / aberration.
124. tyrrhena Stauder, 1915
Original combination. “Coenonympha arcania tyrrhena m. subsp. nov.” Stauder, 1915 Zeit. Wiss. Ins. Biol. 11: 1, is aed MS Si
Current combination. Coenonympha arcania tyr- rhena Stauder, 1915.
Current status. Junior subjective synonym of Co- enonympha arcania arcania (Linnaeus, 1761).
Type material. Syntype 13 (ZMH 827761) (Fig. 124). “Type” // “Calabria / Umg. Paola-Co-/Senza 18.6.1920 / H. Stauder” // “Type / v. tyrrhena Stauder” // “arcania v. / tyrrhena Staud.” // “ZMH 827761”.
Type locality. Italy: Monte Martinello bei 1000 m. [Castello Cabiaglio, Province of Varese, Italy].
125. ab. energica Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova energica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Coenonympha corinna ab. energica Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 1¢ (ZMH 827740) (Fig. 125). “Type / Coen. corinna Hb. / ab. energi- ca Bub. / Otto Bubacek Z.B.G. 1922” //“Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827740”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
126. ab. caeca Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova caeca Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
239
Current combination. Coenonympha corinna ab. caeca Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827741) (Fig. 126). “Type / Coen. corinna Hb. / ab. cae- ca Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827741”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.’’) and is hence unavailable.
127. ab. anophthalmica Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova anophthalmica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Coenonympha corinna ab. anophthalmica Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827742) (Fig. 127). “Type / Coen. corinna Hb. / ab. anophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank la- bel] // “ZMH 827742”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.’) and is hence unavailable.
128. ab. saturata Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova saturata Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Coenonympha corinna ab. saturata Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827743) (Fig. 128). “Type / Coen. corinna Hb. / ab. saturata Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827743”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.’’) and is hence unavailable.
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240 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
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ZMH 827742
Plate 18. 123. Coenonympha arcania f. rufobrunnea Warnecke, 1942; 124. Coenonympha arcania tyrrhena Stauder, 1915; 125. Coenonympha corinna ab. energica Bubacek, 1923; 126. Coenonympha corinna ab. caeca Bubacek, 1923; 127. Coenonympha corinna ab. anophthalmica Bubacek, 1923; 128. Coenonympha corinna ab. saturata Bubacek, 1923; 129. Coenonympha corinna ab. nigricans Bubacek, 1923. a. Dorsal view, b. ventral view.
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Evolutionary Systematics 5 2021, 193-261
129. ab. nigricans Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova nigricans Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Coenonympha corinna ab. nigricans Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827744) (Fig. 129). “Type / Coen. corinna Hb. / ab. nig- ricans Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827744”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
130. ab. macrophthalmica Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova macrophthalmica Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Coenonympha corinna ab. macrophthalmica Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 24'3' (ZMH 827745—827746) (Fig. 130). “Type / Coen. corinna Hb. / ab. macrophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827745”; “Type / Coen. corinna Hb. / ab. macrophthalmica Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juli 1921 / Col. O. Bubacek” // [blank la- bel] // “ZMH 827746”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
131. ab. gynandra Bubacek, 1923
Original combination. “Coenonympha corinna Hb. ab. nova gynandra Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Coenonympha corinna ab. gynandra Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 246 (ZMH 827747—827748) (Fig. 131). “Type / Coen. corinna Hb. / ab. gynandra Bub. / Otto Bubacek Z.B.G. 1922” //
241
“Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827747”; “Type / Coen. corinna Hb. / ab. gynandra Bub. / Otto Bubacek Z.B.G. 1922” // “Evisa Corse / Juni 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827748”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of C. corinna (Hubner, 1804). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.’’) and is hence unavailable.
132. f. pulcherrima Warnecke, 1942
Original combination. “Coenonympha iphis Schiff., 9 n.f. pulcherrima.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102.
Current combination. Coenonympha glycerion f. pulcherrima Warnecke, 1942.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 12 (ZMH 827738) (Fig. 132). “Molln 1. lotz / 30.6.35 [illegible]” /! “30.6.35 / Molln” // “9 n.f. pulcherrima / Warn. Type / Tris 1942, 102” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “Type” // “ZMH 827738”.
Original locality. Germany: “Molln (Lauenburg)*; Schleswig-Holstein, Herzogtum Lauenburg, MOlIn.
Remarks. Warnecke (1942) proposed this name as a form of C. iphis ([Denis & Schiffermuller], 1775). According to article 45.6.1 (ICZN 1999), it is infra- subspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspe- cific entity, and is hence unavailable. From the text it is clear that the author describes an aberration and not a subspecies.
133. ab. oikeia Dannehl, 1927
Original combination. “Coenonympha iphis Schiff. ab. oikeia Dhl.” Dannehl, 1927 Mitt. Minch. Ent. Ges. 17: 5.
Current combination. Coenonympha glycerion ab. oikeia Dannehl, 1927.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 835591) (Fig. 133). “25.6” // “Italia central. / Mti. Sim- bruini/ coll. F Dannehl” // “oikeia / Dhl. / Cotype. / G.” // “Cotype” // “25.6. 1926 / Mti. Simbruini / Mittel.Italien” // “ZMH 835591”.
Original locality. Italy: “Mt. Simbruini von etwa 1400 m Hohe”.
Remarks. Dannehl (1927) proposed this name as an aberration of C. iphis ({Denis & Schiffermiller], 1775). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
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242
134. iranica Schwingenschuss, 1939
Original combination. “Coenonympha leander Esp. nov. ssp. iranica Sch.” Schwingenschuss, 1939 Ent. Z. 53: 62:
Current combination. Coenonympha leander irani- ca Schwingenschuss, 1939.
Current status. Valid subspecies.
Type material. Syntype 14 (ZMH _ 827714) (Fig. 134). “Pers. Elbursgeb. / Kendevanpass / 2800 m, 3.-8.7.36 / Schwingenschuss” // “Cotypus” // “ssp. irani- ca Schwing.” // “Coll. Bytinski-Salz / Eing. Nr. 20 — 60” /1 “ZMH 827714”.
Type locality. Iran: “Am Kendevan (Sch., W., Pf.) von 2600 m bis 3000 m” [Kandovan pass].
135. rhenana Gradl, 1933
Original combination. “Coenonympha oedipus F. sub- species rhenana” Gradl, 1933 Int. Ent. Z. 27: 261.
Current combination. Coenonympha_ oedippus rhenana Gradl, 1933.
Current status. Junior subjective synonym of nomi- notypical Coenonympha oedippus (Fabricius, 1787).
Type material. Syntypes 4¢¢ (ZMH 827722- 827725) (Fig. 135). “Feldk / Vorarlberg / Prof. F. Gradl / 31.7.32 / Rheinau” // “Subsp. / rhenana / F-Gradl” // “Cotype” // “L.23 / Coenonympha / oedippus - | rhena- na Gradl” // “Sig. G. Warnecke / Eing Nr 5, 1949” // “ZMH 827722”; “Vorarlberg / Prof. F. Gradl / Rheinauen /12.7.27 // “Subsp. / rhenana / F-Gradl” // “Cotype” // “ZMH 827723”; “Vorarlberg / Prof. F. Gradl / Feldkirch. Rhein / auen. 12.7.31” // “Subsp. / rhenana / F-Gradl” // “Cotype” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827724”; “Vorarlberg / Prof. F. Gradl / Feldk-Rheinau / 28.7.32” // “Subsp. / rhenana / F-Gradl” /1 “Co-Type” // “Type” // “ZMH 827725”.
Type locality. Germany: “in den Niederungen des Rheintales”. [in the lower region of the Rhine valley].
136. galvagnii Stauder, 1923
Original combination. “C. p. galvagnii m. nova sub- spec. (partim aberr.)” Stauder, 1923 Zeit. Wiss. Ins. Biol. 1259 ty Ser a6 13:
Current combination. Coenonympha pamphilus galvagnii Stauder, 1923.
Current status. Junior subjective synonym of nomi- notypical Coenonympha pamphilus (Linnaeus, 1758).
Type material. Syntype 14 (ZMH 827739) (Fig. 136). “Type” / “H. Stauder / Litorale austr. / Triest Umgbg. / 12/8/1913” // “pamphilus | galvagnii Staud.” // “Type” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827739”.
Type locality. Italy: Bologna “Bei Grado auf den La- gunen in der g. v. und sonst im Kustengebiete, bei Triest, Gorz.” [Bologna, vicinity of Costarotara and Piano As- tore of the Sibillini Mountains, 1200-1600].
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
137. f. maculata Warnecke, 1942
Original combination. “Coenonympha tiphon Rott., °, n. f. maculata.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 103.
Current combination. Coenonympha tullia f. macu- lata Warnecke, 1942.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 229 (ZMH 827762-827763) (Fig. 137). “Bahrenfeld / 1913” // “9 n.f. maculata / Warn. Cotype / Iris 1942, 103” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827762”; “Sege- berg / Juli 13” // “Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827763”.
Original locality. Unclear; Germany: Bahrenfeld, Hamburg “aus Holstein.”; publication states: Holstein, la- bel of cotype specimen from Warnecke states Bahrenfeld, ?, (one further specimen without type label: Segeberg)
Remarks. Warnecke (1942) proposed this name as a form of C. tiphon (Rottemburg, 1775). According to article 45.6.1 (ICZN 1999), it is infrasubspecific if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, and 1s hence unavailable. From the text and already from the title of the paper it is clear that the author describes an aberra- tion. Papilio tiphon Rottemburg, 1775 is junior subjec- tive synonym of Coenonympha tullia (Miller, 1764).
138. flavofasciata Heyne, 1895
Original combination. “Evebia flavofasciata Heyne.” Heyne, [1895]; in Ruhl & Heyne, Die Pal. Gross-Schmett. 1: 805.
Current combination. Erebia flavofasciata Heyne, 1895.
Current status. Valid species.
Type material. Paratype 14 (ZMH 833469) (Fig. 138). “Campolungo / Pass / Oberst v. Nolte” // “Flavofasciata” // “Type oder Paratype / e coll. von Nolte / s. Heyne-Ruhl, 1895, 805, / der keine Samml. hatte. / von Fuchs-Hbg ded.” // “ZMH 833469”.
Type locality. Italy: Tessin, Campolungo pass close to Fusio.
139. peneplana Berger, 1936
Original combination. “Erebia aethiops ESPER, race peneplana, nov. race” Berger, 1936 Lambillionea 36: 199.
Current combination. Erebia aethiops peneplana Berger, 1936.
Current status. Valid subspecies.
Type material. Paratypes 3¢6¢ (ZMH 835588-— 835590) (Fig. 139). “PARATYPE / Collection / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 5/8/1936 / Coll. Luc. BERGER.” // [blank label] // “ZMH835588”; “PARATYPE / Collection / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 5/8/1936 / Coll. Luc. BERGER.”
Evolutionary Systematics 5 2021, 193-261 243
Ly pe Coen.corlana Hb... nba. hance. Bales
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131a*
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| ‘fu fi manasa
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1cm
Plate 19. 130. Coenonympha corinna ab. macrophthalmica Bubacek, 1923; 131. Coenonympha corinna ab. gynandra Bubacek, 1923; 132. Coenonympha glycerion f. pulcherrima Warnecke, 1942; 133. Coenonympha glycerion ab. oikeia Dannehl, 1927; 134. Coenonympha leander iranica Schwingenschuss, 1939; 135. Coenonympha oedippus rhenana Gradl, 1933. a. Dorsal view, b. ventral view.
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244 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
// {blank label] // “ZMH835589”; “PARATYPE / Collec- tion / Luc. Berger” // “Tellin. (R.d. Boyes) / BELGIQUE / 16/8/1934 / Coll. Luc. BERGER.” // [blank label] // “ZMH835590”.
Type locality. Belgium: “env. de Han-sur-Lesse”’.
140. clorinda Hartig, 1940
Original combination. “Erebia epiphron Fn. ssp. clorin- da n.” Hartig, 1940 Mem. Soc. Ent. Ital. 18: 186.
Current combination. Erebia epiphron clorinda Hartig, 1940.
Current status. Junior subjective synonym of Erebia epiphron amplevittata Verity, 1921.
Type material. Paratypes 1619 (ZMH 827718- 827719) (Fig. 140). “Paratypus 9 / Er. epiphr. / clorinda / Hartig” // “1700 m / Marche / Bolognola / 20.VI 1937 / Querci” // [blank label] // “ZMH 827718”; “Paratypus 9 / Er. epiphr. / clorinda / Hartig” // “1700 m / Marche / Bolognola / 20. VI 1937 / Querci” // [blank label | // “ZMH 827718”; “Paratypus ¢ / Er. epiphr. / clorinda Hartig” // “1700 m / Marche / Bolognola / 20. VI 1937 / Querc1” // “Tblank label] // “ZMH 827719”.
Type locality. Italy: “Bolognola”.
141. vogesiaca Goltz, 1914
Original combination. “Erebia epiphron vogesiaca” Goltz, 1914 Dt. Ent. Z. Iris 28:107—109.
Current combination. Erebia epiphron vogesiaca Goltz, 1914.
Current status. Junior subjective synonym of Erebia epiphron mackeri Fuchs, 1914).
Type material. Syntype 14 (ZMH 827679) (Fig. 141). “COTYPE / vogesiaca / v.d.Goltz” // “VII 1913 / Faux- kopf / Vogesen” // “Fauxkopf / Juli 1913 / Kesenheimer” /1 “ZMH 827679”.
Type locality. France: “Vogesen”’.
142. leonhardi Fruhstorfer, 1918
Original combination. “Evebia melas leonhardi sub- spec. nova” Fruhstorfer, 1918 Arch. Naturgesch. 84 AP NS 1.
Current combination. Erebia melas leonhardi Fruhstorfer, 1918.
Current status. Valid subspecies.
Type material. Syntypes 11¢9 (ZMH 827727- 827737) (Fig. 142). “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” /1 “ZMH 827727”; “Velebit, Ostaria” // “Cotype” // “Co- typus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827728”, “Velebit, Ostaria” // “Cotype” // “Coty- pus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827729”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH
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827730”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827731”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827732”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827733”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827734”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827735”; “Velebit, Ostaria” // “Cotype” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827736”; “Velebit, Ostaria” // “Cotypus” // “melas leon- hardi Cotype / Velebit Ostria 19[..]” // “Coll. Bytins- ki-Salz / Eing. Nr. 20, 1960” // “ZMH 827737”. Type locality. Croatia: “Velebit, Ostaria”’.
143. paradisi Dannehl, 1929
Original combination. “Erebia stygne O. paradisi Dhl.” Dannehl, 1929 Mitt. Munch. Ent. Ges. 19: 98.
Current combination. Erebia meolans paradisi Dannehl, 1929.
Current status. Junior subjective synonym of nomi- notypical Erebia meolans (de Prunner, 1798).
Type material. Syntypes 3¢¢ (ZMH 835585-— 835587) (Fig. 143). “19.6” / “Apenn. merid. / Mt. Par- adiso / 1500-2000 m / coll. F. Dannehl” // “19.6. 1928 / Mte. Paradiso / Central — Apenninen” // “ZMH 835585”; “22.6” / “Apenn. merid. / Mt. Paradiso / 1500-2000 m / coll. F. Dannehl” // “20.6. 1928 / Mte. Paradiso / Cen- tral — Apenninen” // “ZMH 835586”; “19.6” / “Apenn. merid. / Mt. Paradiso / 1500-2000 m / coll. F. Dannehl” // “19.6. 1928 / Mte. Paradiso / Central — Apenninen” // “ZMH 835587”.
Type locality. Italy: “Monte Paradiso 1600-1900 m, Monte Rotella, Montagna Grande 1800-2100 m (stil. Abruzzen).”
144. benacensis Warren, 1933
Original combination. “Erebia ottomana ssp. benacen- sis nov.” Warren, 1933 Ent. Rec. 45: 40.
Current combination. Erebia ottomana benacensis Warren, 1933.
Current status. Junior subjective synonym of nomi- notypical Erebia ottomana Herrich-Schaffer, (1847).
Type material. Syntypes 5¢¢ (ZMH 833449- 833453) (Fig. 144). “4.7” // “Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dannehl” // “4.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // “ZMH 833449”; “4.7” // “Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dan- nehl” // “4.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // “ZMH 833450”; “5.7” // “Teriolis merid. / Monte Baldo / 2000 m. / coll. F. Dannehl” // “5.7. 1930 / Mte. Baldo 2000 m / (Gardasee)” // “ZMH 833451”.
Type locality. Italy: “Mte. Baldo near Lake Garda”.
Evolutionary Systematics 5 2021, 193-261 245
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ae Ubud. U... Motles | Flavofasciata
Type 0
Tellin. (R.d.Boyes.)_ BELGIQUE. £7. Ff /ABE.
Coll-Lue. BERGER.
ZMH 833469
- Fauxkopf Julh1913 Kesenheimer
ZMH 327679]
ZMH 827719|
140b
141b 1cm
Plate 20. 136. Coenonympha pamphilus galvagnii Stauder, 1923; 137. Coenonympha tullia f. maculata Warnecke, 1942; 138. Erebia flavofasciata Heyne, 1895; 139. Erebia aethiops peneplana Berger, 1936; 140. Erebia epiphron clorinda Hartig, 1940; 141. Erebia epiphron vogesiaca Goltz, 1914. a. Dorsal view, b. ventral view.
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246 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
145. kotzschae Goltz, 1937
Original combination. “Erebia kotzschae” Goltz, 1937 Ent. Rdsch. 54: 363.
Current combination. Paralasa kotzschae (Goltz, 1937).
Current status. Valid species.
Type material. Syntype 146 (ZMH 827715) (Fig. 145). “Nord-Ost-Hindukusch / Nuksan-Pass-Nordseite / Alpenwiesenzone / 3500-4000 m Mitte Juli / leg. H.&E. Kotzsch” // “COTYPE / kotzschae / v.d. Goltz” // “Kots- chi v.d. Goltz” // “Coll. Bytinski-Salz / Eing Nr 20 — 60” /1 “ZMH 827715”.
Type locality. Afghanistan: Hindukush.
146. maxima Bang-Haas, 1933
Original combination. “Satyrus autonoe maxima O. B.- Haas, subsp. nov.” Bang-Haas, 1933 Ent. Z. 47: 98.
Current combination. Hipparchia autonoe maxima (Bang-Haas, 1933).
Current status. Valid subspecies.
Type material. Syntypes 2¢¢ (ZMH 827681- 827682) (Fig. 146). “Kansu mer. / Min-chan / Tanho / 2500 m. Juli” // “Co-Type / O. Bang-Haas” // “L.13 / Sat. autonoé - / maxima / O.B.H.” // “Sammlung / R. Jahnig / Eing Nr. 29, 1949” // “ZMH 827681”; “Kansu mer. or. / Fukiang / Peilingschan sept. / 2500 m. Juli” // “Co- Type / O. Bang-Haas” // “L.14 / Sat. autonoé - / maxima / O.B.H.” // “Sammlung / R. Jahnig / Eing Nr. 29, 1949” // “ZMH 827682”.
Type locality. China: “Kansu mer., Lihsien, Kialing FluB, 2500 m.”
147. persiscana Verity, 1937
Original combination. “Eumenis allionii, G.-H. = fat- ua Freyer, persiscana, nom. nov.” Verity, 1937 Ent. Rec. 49: 100.
Current combination. Hipparchia fatua persiscana (Verity, 1937).
Current status. Valid subspecies.
Type material. Syntypes 31¢¢ (ZMH 827683- 827713) (Fig. 147). “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827683”; “Tran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827684”; “Iran / Keredj 1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827685”; “Iran / Keredj 1600 m/ VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827686”; “Iran / Keredj 1600 m/ VIII 1936 / leg. F. Brandt / coll. By- tinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing.
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Nr. 20 - 60” // “ZMH 827687”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Coty- pus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827688”; “Iran /Keredj 1600 m/ VIII 1936/leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eimg. Nr. 20 - 60” // “ZMH 827689”; “Iran / Keredj 1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827690”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827691”; “Tran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. By- tinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827692”, “Iran / Keredj 1800 m / 21.8.1936 / Brandt” // “Cotypus” // “ssp. persicana / Vrty” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827693”; “Iran / Keredj 1600 m / VHI 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827694”; “Tran / Keredj 1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytins- ki-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827695”; “Iran / Keredj 1800 m / 25/8 1936 / Brandt” //“Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827696”; “Iran / Keredj 1800 m / 25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827697”; “Iran / Keredj 1600 m/ VU 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827698”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827699”; “Tran / Keredj 1800 m / 25/8 1936 / Brandt” // “201” // “Coty- pus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827700”; “Iran / Keredj 1800 m / 21/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827701”; “Iran / Keredj 1200 m / 1936 / Brandt” // “1600 25/VIIT’ // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827702”; “Iran / Kered] 1600 m/ VII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827703”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- tinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827704”; “Tran / Keredj 1600 m/ VHI 1936 / leg. F. Brandt / coll. Bytins- ki-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827705”; “Iran / Keredj (1200 m.) / 1936 / Brandt” //“Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827706”; “Iran / Keredj 1600 m / 25/8 1936 / Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827707”; “Iran / Keredj 1600 m/ VU 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827708”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” //“ZMH 827709”; “Iran / Keredj 1600 m/ VHI 1936 /leg. F. Brandt / coll. Bytins- ki-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827710”; “Iran / Keredj 1200 m/ 1936
Evolutionary Systematics 5 2021, 193-261 247
Apenn, merid. Mt. Paradiso 1500-2000 m
Cotypus
+
|. a ZMH 835585.
erid, aldo
Teriolis
Monte Nord-Ost-Hindukusch
Nuksan-Pass-Nordseite Alpenwiesenzone 3500-4000m Mitte Juli leg. H.&E. Kotzsch
[ou Bytinski-Salz: Eing Nr 20 - 69:
f Sammlung R.Jaltnig Eing.Nr.29,1949
Kansu mer. or. Fukiang , Peilingschan sept. 2500 m. Juli
Plate 21. 142. Erebia melas leonhardi Fruhstorfer, 1918; 143. Erebia meolans paradisi Dannehl, 1929; 144. Erebia ottomana benacensis Warren, 1933; 145. Paralasa kotzschae (Goltz, 1937); 146. Hipparchia autonoe maxima (Bang-Haas, 1933). a. Dorsal view, b. ventral view.
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/ Brandt” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20 - 60” // “ZMH 827711”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. Bytinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827712”; “Iran / Keredj 1600 m / VIII 1936 / leg. F. Brandt / coll. Bytinski-Salz” // “Cotypus” // “Coll. By- tinski-Salz / Eing. Nr. 20, 1960” // “ZMH 827713”.
Type locality. Iran: Tehran, “Keredj [Karaj], in the El- burz Mountains, at m. 1500 to 1600”.
148. ab. pallida Bubacek, 1923
Original combination. “Satyrus neomiris God. ab. nova pallida Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Hipparchia neomiris ab. pal- lida (Bubacek, 1923).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 1¢ (ZMH 827726) (Fig. 148). “Type / Sat. neomiris God / ab. pal- lida Bub. / Otto Bubacek Z.BG. 1922” // “Col de Verg. Cor- / se Juli 1921 / Col. O. Bubacek” // [blank label] // “ZMH 827726”.
Original locality. France: Corse, Col de Verg.
Remarks. Bubacek (1923) proposed this name as an aberration of S. neomiris (Godart, 1824). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infra- subspecific name (the author used “aberration”, “ab.’) and is hence unavailable.
149. ab. albinervata Warnecke, 1944
Original combination. “[Satyrus semele L.] n. ab. albin- ervata” Warnecke, 1944 Dt. Ent. Z. Iris 57: 40.
Current combination. Hipparchia semele ab. albin- ervata (Warnecke, 1944).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Holotype” 19 (ZMH 827749) (Fig. 149). “Insel Sylt / Kampen / VII 1942” // “n.f. albinervata / Warnecke / Type” // “Holo- type” // “Sammlung G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827749”.
Original locality. Germany: “Sylt, Kampen”.
Remarks. Warnecke (1944) proposed this name as a form of S. semele (Linnaeus, 1758). As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspe- cific name (the author used “aberration”, “ab.”) and is hence unavailable.
150. siviae Fuchs, 1954
Original combination. “Lymanopoda shefteli siviae” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Current combination. Lymanopoda shefteli siviae Fuchs, 1954.
Current status. Valid subspecies.
Type material. Lectotype 14 (ZMH 827764) and 3 SS paralectotypes (ZMH 827765—827767) (Fig. 150). “Lymnopoda / siviae / v. Fuchs / Cordillere von / Siv- ia, Ayacucho / 2500 m.” // “Type” // “Holotype / Lym- nopoda siviae / von Fuchs” // “Lymnopoda galactea / shefteli Dyar / G. Lamas. det. 1978” // “Lectotype 3 / Lymnopoda / siviae Fuchs / by G. Lamas, 1978” // “Sammlung / W. v. Fuchs / Eing Nr. 4, 1956” // “ZMH 827764”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // “285 / galactea ab. / Rio Piene 2600 m. 7/20” // “ZMH 82765”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // “ZMH 82766”; “Rio Piene / ca. 2600 m / 7/1920 / Type” // “ZMH 82767”.
Type locality. Peru: “... oberhalb Sivia gefangen, [above Sivia] 2500-3000 m”.
(<4
151. cocuzzana Stauder, 1914
Original combination. “Melanargia arge Sulz. (=am- phitrite Hb.) cocuzzana m., subspec. nova” Stauder, 1914 Zeit. Wiss. Ins. Biol. 10: 375, f. 12-15.
Current combination. Melanargia arge cocuzzana Stauder, 1914.
Current status. Junior subjective synonym of nomi- notypical Melanargia arge (Sulzer, 1776).
Type material. Paratypes 3¢6¢ (ZMH 827673- 827675) (Fig. 151). “Calabria / Umg. Paola-Co-senza 4.6.1913 / H. Stauder” // “arge cocuzzana Stau- /der Paratype” // “ZMH 827673”; “Calabria / Umg. Paola- Co-senza 4.6.1913 / H. Stauder” // “cocuzzana” // “arge cocuzza- / na Stauder / Paratype” // “ZMH 827674”; “Ital. mer. / pen.-Surrentina / 8.6.1913 / H. Stauder” // “Arge cocuzzana / Stauder” // “arge cocuzzana Stau- / der Para- type” // “ZMH 827675”.
Type locality. Italy: vicinity of Paola, Monte Cocuzzo “ an den Hangen [slopes of | des Monte Martinello und Cocuzzo bei Paola”.
152. ab. huenei Hirschke, 1910
Original combination. “Pararge hiera var. ominata Krul. ab. Huenei” Hirschke, 1910 Verh. Zool. Bot. Ges. Wien 60: 411.
Current combination. Lasiommata petropolitana ominata ab. huenei (Hirschke, 1910).
Current status. Infrasubspecific and hence unavail- able name..
Original material. Labelled as “Type” 1¢ (ZMH 827717) (Fig. 152). “Estonia / Lechts / FR. V. HUENE” // “Type / Verh. z.b.g./ LX. 1910” // “Pararge hiera / var. ominata / ab. huenei / Hirschke / Estland” // “Coll. By- tinski-Salz / Eing. Nr. 20 — 60” // “ZMH 827717”.
Original locality. Estonia: “aus Esthland”.
Evolutionary Systematics 5 2021, 193-261 249
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Plate 22. 147. Hipparchia fatua persiscana (Verity, 1937); 148. Hipparchia neomiris ab. pallida (Bubacek, 1923); 149. Hipparchia semele ab. albinervata (Warnecke, 1944); 150. Lymanopoda shefteli siviae Fuchs, 1954; 151. Melanargia arge cocuzzana Stauder,
1914. a. Dorsal view, b. ventral view.
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250
Remarks. Hirschke (1910) proposed this name as an aberration of P. h. ominata Krulikowsky, 1903, therefore, as stated by articles 45.6.1 and 45.6.2 ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable.
153. felix Warnecke, 1929
Original combination. “Pararge felix n. spec. Warn.” Warnecke, 1929 Int. Ent. Z. 22: 365.
Current combination. Lasiommata felix (War- necke, 1929).
Current status. Valid species.
Type material. Paratype 12 (ZMH _ 827721) (Fig. 153). “S.W. Arabia / Yemen, Sanaa / II.1928. Ex- pedi- / tion Rathjens” // “28.2. /Sanaa” // “Pararge felix / Warnecke 1929 / 9 Paratype” // “Para-Type” // “Pararge maera - | felix Warn.” // “Sig. G. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827721”.
Type locality. “Yemen, Sanaa”.
154. ab. diluta Bubacek, 1923
Original combination. “Pararge aegeria L. ab. nova diluta Bub.” Bubacek, 1923 Verh. Zool. Bot. Ges. Wien 72: (30).
Current combination. Pararge aegeria ab. diluta Bubacek, 1923.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 827678) (Fig. 154) “Type / Par. aegeria L. / ab. diluta Bub. / Otto Bubacek ZBG 1922” // [blank label] // “Evisa, Corse / Juli 1921 / Col. O. Bubacek” // “ZMH 827678”.
Original locality. France: Korsika [Corse].
Remarks. Bubacek (1923) proposed this name as an aberration of P. aegeria (Linnaeus, 1758), therefore, as stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable.
155. f. albiplaga Warnecke, 1942
Original combination. “Pararge algeria L. subsp. egerides Stdgr. n. f. albiplaga 9°.” Warnecke, 1942 Dt. Ent. Z. Iris 56: 102.
Current combination. Pararge aegeria egerides f. albiplaga Warnecke, 1942.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 19 (ZMH 827668) (Fig. 155). “Holstein / Segeberg / 2.-10.8.35” // “nf. albiplaga / Warn. Type / Iris 1942, 102” // “Type” // “ZMH 827668”.
Original locality. Germany: Holstein [label indicates Segeberg].
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Remarks. Warnecke (1942) proposed this name as a form of P. a. egerides (Staudinger, 1871), therefore, as stated by article 45.6.1 it is deemed to be an infrasubspe- cific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. In the original descrip- tion the name “aegeria” is wrongly spelled “algeria” (M. Wiemers pers. comm. ).
156. ab. hermini Hirschke, 1910
Original combination. “Pararge megera var. lyssa B. ab. Hermini” Hirschke, 1910 Verh. Zool. Bot. Ges. Wien 60: 411.
Current combination. Lasiommata megera lyssa ab. hermini (Hirschke, 1910).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 1¢ (ZMH 827720) (Fig. 156). “Hirschke.Wien / Zara / Mai” // “Type / Verh. Z.G.B. / LX, 1910” // “Pararge megera / var. Ly- ssa ab. / hermini Hirschke / Dalmatien 5.1909” // “Coll. Bytinski-Salz / Eing Nr 20 — 60” // “ZMH 827720”.
Original locality. Croatia: Dalmatia, vicinity of Gra- vosa and Zara.
Remarks. Hirschke (1910) proposed this name as an aberration of P. m. lyssa “B.” (sic, recte Hubner, 1829), therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infrasubspecific rank) and is hence unavailable.
157. picota Fuchs, 1954
Original combination. “Argyrophorus picota” Fuchs, 1954 in Titschak, Beit. zur Fauna Perus 4: 84.
Current combination. Punargentus angusta picota (Fuchs, 1954).
Current status. Valid subspecies.
Type material. Lectotype 14 (ZMH 827750) (Fig. 157) 7319 paralectotypes (ZMH 827751-827758). “Argyro- phorus / picota 3 / y. Fuchs / Cordillere von / Ayacucho / 4500 m” // “Punargentus lamna / angusta Weymer / det. W. Heinrich 3” // “Type” // “Lectotype ¢ / Argyrophorus / picota Fuchs / by G. Lamas, 1978” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “ZMH 827750”; “Punar- gentus lamna / angusta Weymer / det. W. Heinrich 3” // Argyrophorus | picota 3 / v. Fuchs / Cordillere von / Ay- acucho / 4500 m” // “Type” // “Sammlung / W. v. Fuchs / Eing. Nr. 4, 1956” // “ZMH 827751”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich 3” // Coll. H. Rédinger / Eing. Nr. 30, 52” // ZMH 827752”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich 3” // Coll. H. Rédinger / Eing. Nr. 30, 52” /1 ZMH 827753”, “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punargentus lamna / angusta Weymer / det. W. Heinrich @” // “133 / angusta 8/20 / Ayacucho 4/5000 m”
Evolutionary Systematics 5 2021, 193-261 251
LW, brabs ies ina
1914. Et pede~ Him Fates cus
Porarce hiera Var. oa . n Ve ‘hutnei Est lineal’ rsthiee
Coll, Bytingki-Salz: ‘Eing, Nr., 20 - 69:
Beosgextislenme nc STE Weyer
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Baring i'ma almatien F.71999
157b
11cm
Plate 23. 152. Lasiommata petropolitana ominata ab. huenei (Hirschke, 1910); 153. Lasiommata felix (Warnecke, 1929); 154. Pararge aegeria ab. diluta Bubacek, 1923; 155. Pararge aegeria egerides f. albiplaga Warnecke, 1942; 156. Lasiommata megera lyssa ab. hermini (Hirschke, 1910); 157. Punargentus angusta picota (Fuchs, 1954). a. Dorsal view, b. ventral view.
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252) Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
// “Coll. H. Rodinger / Eing. Nr. 30, 52” // ZMH 827754”: “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punar- gentus lamna / angusta Weymer / det. W. Heinrich 3” // Coll. H. Rédinger / Eing. Nr. 30, 52” // ZMH 827756”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punar- gentus lamna / angusta Weymer / det. W. Heinrich 3” // Coll. H. Rédinger / Eing. Nr. 30, 52” // ZMH 827757”; “Ica / Ayacucho / ca. 4000 m. / 7/1920 / Type” // “Punar- gentus lamna / angusta Weymer / det. W. Heinrich 9” // Coll. H. Rodinger / Eing. Nr. 30, 52” // ZMH 827758”.
Type locality. Peru: near Ayacucho “... Kodillere bei Ayacucho, nur in Hohen tiber 4000 m, bis 5000 m” [Cor- dillere close to Ayacucho, only at altitudes above 4000 m, to 5000 m].
158. arminii Stauder, 1916
Original combination. “Epinephele ida arminii subsp. nova.” Stauder, 1916 Zeit. Wiss. Ins. Biol. 12: 62.
Current combination. Pyronia cecilia arminii (Stauder, 1916).
Current status. Junior subjective synonym of Pyronia cecilia cecilia (Vallantin, 1894).
Type material. Syntype 14 (ZMH _ 827716) (Fig. 158). “Type” // “Calabria mer. / Aspromonte 1600 m. / 26.6.1920 / H. Stauder legit.” // “Ida Arminii / Staud- er” // “Type” // “Sammlung / G. Warnecke / Eing. Nr. 5, 1949” // “ZMH Nr. 5, 1949”,
Type locality. Italy: Calabria.
159. liupiuschani Bang-Haas, 1933
Original combination. “Satyrus actaea liupiuschani O. B.-Haas, subsp. nov.” Bang-Haas, 1933 Ent. Z. 47: 99.
Current combination. Satyrus ferula liupiuschani Bang-Haas, 1933.
Current status. Junior subjective synonym of Satyrus ferula ganssuensis Grum-Grshimailo, 1893.
Type material. Syntype 14 (ZMH 827669) (Fig. 159). “Kansu mer. / Tsing-schui / Liu-pin-schan / 2000 m. Juli” // “Co-Type / O. Bang-Haas” // “L.17 / Sat. ac- taea - / liupiuschani O.B.H.” // “Sammlung / R. Jahnig / Eing. Nr.29,1949” // “ZMH 827669”.
Type locality. China: “Kansu mer., Tsingschui, Liu Piu Schan, 2000 m”.
160. kintschouensis Bang-Haas, 1939
Original combination. “7riphysa phryne kintschouensis O. Bang-Haas, subsp. nov.” Bang-Haas, 1939 Dt. Ent. Z. Ins S3"53"
Current combination. Coenonympha phryne kint- schouensis (Bang-Haas, 1939).
Current status. Junior subjective synonym of Co- enonympha dohrnii nervosa Motschulsky, 1866.
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Type material. Syntypes 2¢¢ (ZMH 827759- 827760) (Fig. 160). “18.” // “Mandschuria mer. occ. / Prov. Fengtien / Kintschou / 100 m. April” // “Co-Type / O. Bang-Haas” // “Phryne / v. kintschouensis / O.B.H.” // “L.21 / Triphysa phyrne - / kintschouensis / O.B.- Haas” // “Sammlung R. Jahnig / Eing Nr. 29, 1949” // “ZMH 827759”; “17.” // “Mandschuria mer. occ. / Prov. Fengtien / Kintschou // “Co-Type / O. Bang-Haas” // “L,.22 / Triphysa phyrne - / kintschouensis / O.B.-Haas” // “Sammlung R. Jahnig / Eing Nr. 29, 1949” // “ZMH 827760”.
Type locality. China: “Manchuria merid., Prov. Feng- tien, Kintschou, 100 m.”.
Unpublished, unavailable names
161. ab. minor Meinhard
Original combination. Melanargia galathea v. sakaria ab. minor Meinhard.
Current status. “in litteris’ name and hence not available.
Original material. Labelled as “Type” 1¢ (ZMH 835597) (Fig. 161). “Type” // 7/7 1920 / Gottingen / Warteberg” // “ZMH 835597”.
Remarks. This taxon cannot be found in the literature. As stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”, or the author expressly gave it infra- subspecific rank) and is hence unavailable.
162. ab. caeca Dannehl
Original combination. Epinephele ida ab. caeca Dan- nehl.
Current status. “in litteris’ name and hence not available.
Original material. Labelled as “Cotype” 14 (ZMH 833467) (Fig. 162). “1.6” // Sicilia / Mt. Cuccio / coll. Dannehl” // “Cotype / caeca / Dhl” // “Cotype.” // “1.6 1932 / Mt. Cuccio, / -Sizilien-” // “ZMH833467”.
Remarks. This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberration”, “ab.”) and is hence unavailable. The current name of the species 1s Pyronia cecilia (Vallantin, 1894).
163. achaltekkensis Holik
Original combination. Satyrus actaea achaltekkensis Holik.
Current status. “in litteris’ name and hence not available.
Original material. Labelled as “Cotypes” 24'3' (ZMH 827670—-827671) (Fig. 163). “Transcaspia / Jablonowka / Achal Tekke / 2000 m. Juli” // “Co-Type” // “L.18 / Sat.
Evolutionary Systematics 5 2021, 193-261 253
Kansu mer- Tsing-schui ‘Liu-pin-schan! 2000 m. Juli:
158a
' Sammlung : RJahhig | |Eing.Nr.29,1949 |
158b
Mandschuria mer.oce, rov. Fengtien
Kintgthou (100m ll
“Sammlung RJabnig 5 Fina Nr.29,1949 :
> = <= 2MH_835597
Transcaspia Jablonowka Achal Tekke 2000m, Juli
Plate 24. 158. Pyronia cecilia arminii (Stauder, 1916); 159. Satvrus ferula liupiuschani Bang-Haas, 1933; 160. Coenonympha phryne kintschouensis (Bang-Haas, 1939); 161. Melanargia galathea v. sakaria ab. minor Meinhard; 162. Epinephele ida ab. caeca Dannehl; 163. Satyrus actaea achaltekkensis Holik. a. Dorsal view, b. ventral view.
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actaea - | achaltekkensis Hol.” // “Sammlung / R. Jahnig / Eing. Nr. 29, 1949” // “ZMH 827670”; “Transcaspia / Jablonowka / Achal Tekke / 2000 m. Juli” // “Co-Type” // “L.19/ Sat. actaea - / achaltekkensis Hol.” // “Sammlung /R. Jahnig / Eing. Nr. 29, 1949” // “ZMH 827671”.
Original locality. Turkmenistan, Achal Tekke.
Remarks. This taxon cannot be found in the litera- ture. The current name is Satyrus iranica daubi Gross & Ebett 1975:
164. rossarmenia Bang-Haas
Original combination. Satyrus actaea rossarmenia Bang-Haas.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Cotype” 14 (ZMH 827672) (Fig. 164). “Kagysman / Russ. Armen” // “Co- Type” //“L.16 / Sat. actaea / rossarmenia ?” // “Sammlu- ng R. Jahnig / Eing. Nr.29, 1949” // “ZMH 827672”.
Remarks. This taxon cannot be found in the literature. The current name is Satyrus amasina Staudinger, 1861.
Family Riodinidae Grote, 1895 Subfamily Nemeobiinae Bates 1868 Unpublished, unavailable names
165. ab. friesi Meinhard
Original combination. Hamearis lucina ab. friesi Meinhard.
Current status. “in litteris” name and hence not available.
Original material. Labelled as “Type” 14 (ZMH 835594) (Fig. 165). “Type” // “ab. friesi Mein.” // “10/5 1911 / Gottingen / Hainberg” // “ZMH 835594”.
Remarks. This taxon cannot be found in the literature. As stated by article 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecific name (the author used “aberra- tion’, “ab.”) and is hence unavailable. The current name is Hamearis lucina (Linnaeus, 1758).
Family Hesperiidae Latreille, 1809 Subfamily Pyrginae Burmeister, 1878 Tribe Carcharodini Verity, 1940
166. wissmanni Warnecke, 1934
Original combination. “Carcharodus swinhoei Wats. nov. subsp. Wissmanni Warn.” Warnecke, 1934 Mitt. Munch. Ent. Ges. 24: 20.
Current combination. manni Warnecke, 1934.
Current status. Valid subspecies.
Type material. Paratypes 5<¢'¢' (ZMH 827637-ZMH 827638, ZMH 827640—-ZMH 827642) (Fig. 166). “S. W. Arabien / Sana’a (Yemen) / 27.2.28 v. Wissm.” // “Sig. G.
Carcharodus alceae wiss-
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Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
Warnecke / Eing. Nr. 5, 1949” // “Carcharodus | Swin- hoei Wats. / subsp. wissmanni / Warn. / Cotype” // “Coll. Warnecke / Eing. Nr. 5, 1949” //“ZMH 827637”; “Alceae Swinhoei Wats. / S.W. Arabia / Sana’a (Yemen) 27.2.28 / Rathjens - v. Wissmann” // “Carcharodus | Swinhoei Wats. / subsp. wissmanni / Warn. / Cotype” // “Coll. War- necke / Eing. Nr. 5, 1949” // “ZMH 827638”, “Stid-Ara- bien / Yemen: San’aa / 8.8.31 Rathjens” // “Paratype” // Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827640”; “Stdwestarabien / Yemen: San’aa / Rathjens 24.6.31” // “Paratype” // Coll. Warnecke / Eing. Nr. 5, 1949” // “ZMH 827641”; “Stidwestarabien / Yemen: San’aa / Rathjens 24.7.31” // “Paratype” // “ZMH 827642”.
Type locality. Yemen: Sanaa “Stdwest-Arabien, Sa- na’a (Yemen)”
167. v. postesseloides Verity, 1938
Original combination. “Spialia orbifer, Hub., race tes- selloides, H.-S., II gen. postesseloides, nom. nov.” Verity, 1938 Ent. Rec. 50 Suppl. (4).
Current combination. Spialia orbifer orbifer v. post- esseloides Verity, 1938.
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 443 (ZMH 833498—-833501) (Fig. 167). “postesseloides Vrty” // “Macedonia / Olympus, 2500’ / Aug 24, 1935 / Romei” // [blank label] // “ZMH 833498”; “postesseloides Vrty” // “Macedonia / Olympus, 2500’ / July 25, 1936 / Rome” // [blank label] // “ZMH 833499”; “postesseloides Vrty” // “Macedonia / Olympus, 2500’ / July 6, 1936 / Romei” // [blank label] // “ZMH 833500”; “postesseloides Vrty” // “Macedonia / Olympus, 2000’ / July 8, 1936 / Romei” // [blank label] // “ZMH 833501”.
Original locality. Greece: vicinity of Skala.
Remarks. Verity (1938) proposed this name as “nom. nov.“ of S. orbifer tesselloides (Herrich-Schaffer, 1845), therefore, as stated by article 45.6.1 it is deemed to be an infrasubspecific name (the author expressly gave it infrasubspecific rank) and is hence unavailable. Spialia orbifer tesselloides is currently regarded as a junior sub- jective synonym of the nominate subspecies.
Tribe Pyrgini Burmeister, 1878
168. centralitaliae Verity, 1920
Original combination. “Hesperia alveus, Hub., race centralitaliae, mihi” Verity, 1920 Ent. Rec. 32: 4.
Current combination. Pyrgus alveus centralitaliae (Verity, 1920).
Current status. Valid subspecies.
Original material. Labelled as “Type” 433 (ZMH 833502—833505) (Fig. 168). “centralitaliae Vrty” // “Marche / Bolognola / 6.VII.1937 / Quer- ci” // [blank label] // “ZMH 833502”; “centralitaliae
Evolutionary Systematics 5 2021, 193-261
Vrty” // “Marche / Bolognola / 13.VII.1937 / Quer- ci” // [blank label] // “ZMH 833503”; “centralitaliae Vrty” // “Marche / Bolognola / 22. VI.1937 / Querci” // [blank label] // “ZMH 833504”; “centralitaliae Vrty” // “Marche / Bolognola / 27.VI.1937 / Querci” // [blank label] // “ZMH 833505”.
Original locality. Italy: Bologna.
Remarks. Verity (1920) proposed this name as a “race” of H. alveus Hiibner, [1803]. As it was given in trinomy and the content of the description clearly indi- cates that a geographical race was meant, this name 1s available as subspecific (article 45.6 ICZN 1999). The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collect- ed from the same geographical location from which the type specimen was collected is called topotype. Accord- ing to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non- type” specimens
169. badachschana Alberti, 1939
Original combination. “Hesperia badachschana nov. sp.” Alberti, 1939 Ent. Rdsch. 56: 107.
Current combination. Pyrgus badachschana (AI- berti, 1939).
Current status. Valid species.
Type material. Holotype 1 ¢ (ZMH 827639) (Fig. 169). “HOLOTYPE < / Hesperia / badachschana” |/ “Bad- achschan / Sebal..|-Tal / Alpenwiesenzone / 2800-3000 m Mitte Juni / leg. H.&E. Kotzsch” // “Coll. Bytinski-Salz Coll / Eing. Nr. 20, 1960 Eing” // “ZMH 827639”.
Type locality. Afghanistan: Badakhshan.
Remarks. We follow SEL declaration on gender agreement (Sommerer 2002), as a result it should stay as badachschana.
170. picena Verity, 1920
Original combination. “Hesperia foulquieri, Obth., race picena, mihi” Verity, 1920 Ent. Rec. 32: 4.
Current combination. Pyrgus foulquieri picena (Verity, 1920).
Current status. Valid subspecies.
Original material. Labelled as “Type” 44'3' (ZMH 833506—833509) (Fig. 170). “Picena Vrty” // “Marche / Bolognola / 2 VII 1937 / Querci” // [blank label] // “ZMH 833506”; “Picena Vrty” // “Marche / Bolognola / 8 Ag 1937 / Querci” // [blank label] // “ZMH 833507”; “Pi- cena Vrty” // “Marche / Bolognola / 8 Ag 1937 / Quer- ci” // [blank label] // “ZMH 833508”; “Picena Vrty” //
255
“Marche / Bolognola / 12 Ag 1937 / Querci” // [blank label] // “ZMH 833509”.
Original locality. Italy: Bolognola, Sibillini Mts.
Remarks. Verity (1920) proposed this name as a “race” of H. foulquieri Oberthur, 1910. As it was given in trinomy and the content of the description clearly in- dicates that a geographical race was meant, this name 1s available as subspecific (article 45.6 ICZN 1999). The year on the specimen labels (1937) indicate specimens were collected 17 years after description (1920) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as “types” is erroneous. The specimen collect- ed from the same geographical location from which the type specimen was collected is called topotype. Accord- ing to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these four specimens are erroneously labelled as types, should be annotated as “Not a Type” and are treated here as “Non- type” specimens.
171. ab. alboradiata Bubacek, 1926
Original combination. “Hesperia sidae Esp. var. on- opordi H.-S. ab. nova alboradiata Bub.” Bubacek, 1926 Verh. Zool. Bot. Ges. Wien 76: 174.
Current combination. Pyrgus sidae ab. alboradiata (Bubacek, 1926).
Current status. Infrasubspecific and hence unavail- able name.
Original material. Labelled as “Type” 14 (ZMH 833497) (Fig. 171). “Type / H. sidae v. onopordi / ab. alboradiata Bub. / Z. B. G. 1926 O. Bubacek” // “Ttalia / Alassio — Liguria / 7/6 1925 / Coll. O. Bubacek” // [blank label] // “ZMH 833497”.
Original locality. Italy: “Alassio, Liguria”.
Remarks. Bubacek (1926) proposed this name as an aberration of H. sidae v. onopordi “H.-S.” (sic, recte Rambur, 1839), therefore, as stated by articles 45.6.1 and 45.6.2 (ICZN 1999) it is deemed to be an infrasubspecif- ic name (the author used “aberration”, “ab.”, or the au- thor expressly gave it infrasubspecific rank) and is hence unavailable. P onopordi Rambur is a distinct species from P. sidae Esper, and both species occur sympatrical- ly in Liguria.
Discussion
With more than 2,000 type specimens of butterflies and moths (Lepidoptera), the Zoological Museum of Ham- burg (ZMH) is one of the most important natural histo- ry archives in Germany. In this catalogue, a total of 414 specimens labelled as “types” belonging to the superfam- ily Papilionoidea are listed and discussed. This includes 23 specimens of 16 “in litteris” (= unavailable) names, 43 topotypes (non-type specimens), 97 original specimens
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256 Reza Zahiri et al.: Lepidoptera type catalogue of Zoological Museum Hamburg: Papilionoidea
/ Pe bP ecccta/ 22. Baty. Vira.
| “Warnacka
faran
5 ce Lene ZMH 833500
es
> Coll. Warnecke | ZEing. Nr. 5, 1949
Coll Bytinski-SalzColl Eing. Nr. 20, 1960Eing.
Plate 25. 164. Satyrus actaea rossarmenia Bang-Haas; 165. Hamearis lucina ab. friesi Meinhard; 166. Carcharodus alceae wissmanni Warnecke, 1934; 167. Spialia orbifer orbifer v.